Ecological dynamics
The Claypan 12-16 ecological site is dominated by Bluebunch wheatgrass, Idaho fescue and little sagebrush. Subdominant species include Sandberg bluegrass, bottlebrush squirreltail, Nevada bluegrass, thickspike wheatgrass, Thurber’s needlegrass, arrowleaf balsamroot, Hooker’s balsamroot and longleaf phlox. Total annual production is 650 lbs/acre in a normal year, 950 lbs/acre in a favorable year, and 350 lbs/acre in a unfavorable year. Structurally, cool season deep-rooted perennial bunchgrasses are dominant, followed by medium height shrubs and perennial forbs. This site is visually dominated by bunch grasses and little sagebrush. Composition by weight is approximately 40-60% grasses, 15-25% forbs and 25-35% shrubs.
Herbivory has historically occurred on the site at low levels of utilization. Native herbivores include pronghorn antelope, mule deer, sage grouse, lagomorphs and rodents. Livestock grazing has become prevalent across this site. Overutilization of resources due to grazing (from livestock and wildlife) can degrade the site and decrease forage availability and quality. This will lead to a decrease in perennial bunch grasses and an increase of invasive species (Williamson, 2020). Annual and perennial invasive species compete with desirable plants for moisture and nutrients.
Wildfire frequency across this site has historically been low. Sagebrush evolved with low intensity wildfire that left a mosaic of burned and unburned patches (Baker, 2006). Annual invasive species such as cheatgrass and medusahead can be troublesome invaders on this site after wildfire, preventing perennial grass and shrub re-establishment. Invasive, annual plant communities increase wildfire frequency and intensity (K. Haubensak, 2009). This could cause the dominate shrub population to shift away from little sagebrush to a shrub population with quicker establishment. A complete absence of wildfire could lead this site to become dominated by Utah Juniper.
High annual precipitation will increase the total plant production. Higher wildfire frequency following annual plant production can be expected due to a larger fuel load (Pilliod, 2017). Extended drought reduces the vigor of perennial grasses and shrubs, while extreme drought may cause plant mortality. Runoff potential following large precipitation events is low if plant communities are present on the landscape. Decreased infiltration, increased runoff, and increased erosion often occur when little sagebrush is removed by frequent wildfires (C.J. Williams, 2018).
State 1
Reference State
The Reference State is a representative of the natural range of variability under pristine conditions. State dynamics are maintained by interactions between climatic patterns and disturbance regimes. Negative feedbacks enhance ecosystem resilience and contribute to the stability of the state. These include the presence of all structural and functional groups, low fine fuel loads, and retention of organic matter and nutrients. Plant community phase changes are primarily driven by fire, periodic drought and/or insect or disease attack.
Community 1.1
This community phase is characteristic of a mid-seral plant community and is dominated by little sagebrush, bluebunch wheatgrass and Idaho fescue. Thurber's needlegrass, Nevada bluegrass, antelope bitterbrush and rabbitbrush are also common on this site.
Potential vegetative composition by weight is about 50 percent grasses, 20 percent forbs and 30 percent shrubs. Total vegetative cover averages 40 to 70 percent and litter cover averages 20 percent.
Table 5. Annual production by plant type
Plant type |
Low (kg/hectare) |
Representative value (kg/hectare) |
High (kg/hectare) |
Grass/Grasslike |
196 |
364 |
532 |
Shrub/Vine |
118 |
219 |
319 |
Forb |
78 |
146 |
213 |
Total |
392 |
729 |
1064 |
Community 1.2
This community phase is characterized by a post-disturbance, early seral, plant community. Little sagebrush and other shrubs are reduced, or patchy. Perennial bunchgrasses and forbs dominate the visual aspect of the plant community.
Disturbance tolerant shrubs such as rabbitbrush and antelope bitterbrush will sprout from the root-crown following low and medium intensity wildfire and may begin to dominate the plant community 2 to 5 years post-disturbance.
Community 1.3
Absence of disturbance allows little sagebrush to mature and dominate the plant community. Perennial bunchgrasses and forbs are reduced in both vigor and productivity due to competition for light, moisture and nutrient resources.
Juniper may also be increasing in cover and number of individual trees. Additional field work is need to determine the extent of juniper on this ecological site and determine if correlation to a more appropriate site is warranted.
Pathway 1.1a
Community 1.1 to 1.2
Wildfire.
Low severity fire creates sagebrush/grass mosaic; higher intensity fires significantly reduce sagebrush cover and lead to early seral community dominated by grasses and forbs.
Frequency and intensity of wildfire is primarily driven by cover and amount of herbaceous vegetation. Under pre-Eurosettlement conditions fire return interval is estimated to be between 20 and 50 years.
Pathway 1.1b
Community 1.1 to 1.3
Time, absence of disturbance and natural regeneration over time allows little sagebrush to dominate site resources. This community phase pathway may be coupled with drought and/or herbivory further reducing herbaceous understory.
Pathway 1.2a
Community 1.2 to 1.1
Time, absence of disturbance and natural regeneration over time allows little sagebrush to recover. Recovery of sagebrush depends on the availability of a local seed source (patches of mature shrubs) as well as precipitation patterns favorable for germination and seedling recruitment. Sagebrush seedlings are susceptible to less than favorable conditions for several years. Completion of this community phase pathways may take decades.
Pathway 1.3a
Community 1.3 to 1.1
Low intensity, patchy wildfire or an aroga moth infestation would reduce little sagebrush overstory creating a mosaic on the landscape. Perennial bunchgrasses and forbs dominate disturbed patches due to an increase in light, moisture and nutrient resources.
Pathway 1.3b
Community 1.3 to 1.2
Wide spread wildfire removes little sagebrush and allows perennial bunchgrasses and forbs to dominate.
State 2
Current Potential
This state is similar to the Reference State 1.0. Ecological function has not changed, however the resiliency of the state has been reduced by the presence of invasive weeds. This state has the same three general community phases. These non-natives can be highly flammable, and can promote fire where historically fire had been infrequent. Negative feedbacks enhance ecosystem resilience and contribute to the stability of the state. These include the presence of all structural and functional groups, low fine fuel loads and retention of organic matter and nutrients. Positive feedbacks decrease ecosystem resilience and stability of the state. These include the non-natives’ high seed output, persistent seed bank, rapid growth rate, ability to cross pollinate and adaptations for seed dispersal.
Management would be to maintain high diversity of desired species to promote organic matter inputs and prevent the dispersal and seed production of the non-native invasive species.
Community 2.1
This community phase is similar to the Reference State Community Phase 1.1, with the presence of non-native species in trace amounts.
Resilience management. The presence of non-native annuals has reduced site resilience. Management actions should focus on maintaining the presence of all functional and structural groups and minimizing wildfire and soil disturbing practices.
Community 2.2
This community phase is characteristic of a post-disturbance, early seral community where annual non-native species are present. Perennial bunchgrasses and forbs recover rapidly following wildfire. Annual non-native species are stable or increasing within the community. Disturbance tolerant shrubs typically recover 2 to 5 years post fire and may dominate the sites for many years.
Community 2.3
This community phase is characterized by decadent sagebrush, reduced perennial bunchgrass and increasing bare ground. Annual non-natives species are stable or increasing due to lack of competition from perennial bunchgrasses. Sandberg bluegrass may increase and become co-dominate with remaining deep-rooted bunchgrasses.
Juniper may also be increasing in cover and number of individual trees. Additional field work is need to determine the extent of juniper on this ecological site and determine if correlation to a more appropriate site is warranted.
Resilience management. This community is at risk of crossing a threshold to another state. This site is susceptible to further degradation from poor grazing management, drought, and/or fire.
Pathway 2.1a
Community 2.1 to 2.2
Fire reduces the shrub overstory and allows for perennial bunchgrasses to dominate the site. Fire may be patchy resulting in a mosaic pattern with patches of mature sagebrush remaining. Annual non-native species are likely to increase after fire.
Pathway 2.1b
Community 2.1 to 2.3
Time and lack of disturbance allows for sagebrush to increase and become decadent. Mature sagebrush is controlling the spatial and temporal distribution of moisture, nutrient and light resources. Native perennial bunchgrasses are reduced due to competition for these resources. Non-native annuals are stable to increasing.
Pathway 2.1a
Community 2.2 to 2.1
Time, lack of disturbance and natural regeneration of sagebrush. The establishment of little sagebrush depends on presence of seed source and favorable weather patterns. It may take decades for sagebrush to recover to pre-disturbance levels.
Pathway 2.3a
Community 2.3 to 2.1
Low intensity wildfire, aroga moth infestation, or brush management with minimal soil disturbance reduces sagebrush overstory and releases herbaceous understory.
Context dependence. Annual non-native species are present and may increase following disturbance.
Pathway 2.3b
Community 2.3 to 2.2
Fire reduces or eliminates the overstory of sagebrush and allows for the understory perennial grasses and forbs to increase. Annual non-native species respond well to fire and may increase post-burn.
State 3
Annual State
Annual non-natives dominated site productivity and site resources. The dominance of non-native annuals control the spatial and temporal distribution of soil moisture, soil nutrients and energy resources. Remaining patches of sagebrush and/or perennial bunchgrass suffer from increased competition and narrowed fire return intervals.
Characteristics and indicators. This state experiences frequent fire due to increased cover and continuity of fine fuels. Fire is frequent enough to prevent the recovery of long-lived native perennials like mountain big sagebrush. Disturbance tolerant shrubs may be present or increasing depending on time since disturbance.
Community 3.1
This community phase in dominated by annual non-native plants such as medusahead or cheatgrass and shallow-rooted perennial grasses like Sandberg bluegrass. Sprouting shrubs such as rabbitbrush may also common. Patches of mature sagebrush may or may not be present.
Community 3.2
This community phase is characteristic of a post-wildfire community where annual non-natives are controlling site resources. Depending on season and/or intensity of fire the visually aspect of the site in dominated annual non-natives and bare ground. Site may be experiencing soil loss.
Resilience management. This community phases is high susceptible to frequent and repeated wildfire. Best management practices prevent sites from reaching this community phase. Management options are extremely limited.
Pathway 3.1a
Community 3.1 to 3.2
Fire reduces or eliminates the overstory shrubs and shallow-rooted perennials and allows for annual non-natives to increase
Pathway 3.2a
Community 3.2 to 3.1
Time and lack of fire allows for sagebrush/rabbitbrush to establish. Probability of sagebrush establishment is very unlikely and dependent on a near-by seed source from unburned patches of sagebrush.
State 4
Tree State
This state is characterized by a dominance of Utah juniper. Little sagebrush and perennial bunchgrasses may still be present, but they are no longer controlling site resources. Soil moisture, soil nutrients and soil organic matter distribution and nutrient cycling have been spatially and temporally altered.
This state is relatively stable due to rapid growth rate and long life span of juniper.
Characteristics and indicators. Juniper is very drought tolerant and has the ability use moisture and nutrients from a wide variety of soil depths, due to its extensive root system. Native species experience reduced productivity and reproductive capacity due to shading and competition for soil moisture and nutrients.
Community 4.1
Juniper dominates overstory and site resources. Trees are actively growing and seedlings may be present. The shrub and grass understory is reduced. Sagebrush is stressed and dying. Trace amounts Sandberg bluegrass and forbs may be found in the interspaces. Annual non-native species are present under tree canopies. Bare ground areas are large and connected.
Transition T1
State 1 to 2
Trigger: Introduction of annual non-native species
Slow variable: Over time the annual non-native plants increase within the community.
Threshold: Any amount of introduced non-native species causes an immediate decrease in the resilience of the site. Annual non-native species cannot be easily removed from the system and have the potential to significantly alter disturbance regimes from their historic range of variation.
Transition T3
State 2 to 3
Trigger: Repeated, widespread and severe fire.
Slow variables: Increased production and cover of non-native annual species over time.
Threshold: Loss of deep-rooted perennial bunchgrasses and shrubs truncates, spatially and temporally, nutrient capture and cycling within the community.
Constraints to recovery. Increased, continuous fine fuels from annual non-native plants modify the fire regime by changing intensity, size and spatial variability of fires. This increased disturbance prevents recovery of long-lived native perennials.
Transition T2
State 2 to 4
Trigger: Presence of juniper
Slow variables: Encroachment of juniper is primarily driven by lack of fire. This may also be coupled with prolonged drought and poor grazing management.
Threshold: Juniper is now controlling energy, moisture and nutrient resources Dominance of juniper results in decreased infiltration and increased runoff, reducing soil moisture and nutrient cycling. Sagebrush and perennial bunchgrass are reduced both vigor and reproductive capacity.
Restoration pathway R2
State 3 to 2
Seeding with native species followed by prescribed grazing
Minimize soil disturbance and maximize non-native annual plant biomass removal during early spring. Combine prescribed grazing with seeding of native species. Continue to protect site from wildfire. Probability of success is extremely low.
Restoration pathway R1
State 4 to 2
Brush management/tree removal with minimal soil disturbance, coupled with seeding of native species. Probability of success very low.
Transition T4
State 4 to 3
Trigger: Catastrophic fire causing a stand replacing event. Or a failed restoration attempt including inappropriate tree removal or rangeland seeding using soil disturbing practices.
Slow variables: Increased production and cover of non-native annual species under tree canopies.
Threshold: Closed tree canopy with non-native annual species in the understory changes the intensity, size and spatial variability of wildfires. Changes in community composition are driven by temporal changes in energy capture, soil moisture and nutrient cycling and result in the loss of perennial bunchgrasses and sagebrush.