Ecological dynamics
Potential vegetation on this site is dominated by mid cool-season perennial grasses. Other significant vegetation includes Gardner saltbush, birdfoot sagebrush, and a variety of forbs. The expected potential composition for this site is about 70% grasses, 10% forbs and 20% woody plants. The composition and production will vary naturally due to historical use, fluctuating precipitation and fire frequency.
As this site deteriorates, species such as blue grama, birdfoot sagebrush and big sagebrush will increase. Plains pricklypear and weedy annuals will invade. Cool season grasses such as rhizomatous wheatgrasses, bottlebrush squirreltail, and Indian ricegrass will decrease in frequency and production.
The Historic Climax Plant Community (description follows the plant community diagram) has been determined by study of rangeland relic areas, or areas protected from excessive disturbance. Trends in plant communities going from heavily grazed areas to lightly grazed areas, seasonal use pastures, and historical accounts have also been used.
The following is a State and Transition Model Diagram that illustrates the common plant communities (states) that can occur on the site and the transitions between these communities. The ecological processes will be discussed in more detail in the plant community narratives following the diagram.
State 1
Rhizomatous Wheatgrasses/Indian Ricegrass Plant Community
Community 1.1
Rhizomatous Wheatgrasses/Indian Ricegrass Plant Community
The interpretive plant community for this site is the Historic Climax Plant Community. This state evolved with grazing by large herbivores, soil less than 15 inches, and periodic fires. The cyclical nature of the fire regime in this community and the shallow soils prevented big sagebrush from being the dominant landscape. This state is comprised of mostly cool season mid-grasses and a variety of forbs and woody species. Potential vegetation is about 70% grasses or grass-like plants, 10% forbs, and 20% woody plants.
The major grasses include rhizomatous wheatgrasses, Indian ricegrass, bottlebrush squirreltail, and bluebunch wheatgrass. Other grasses occurring on the state may include Sandberg bluegrass, blue grama, and prairie junegrass. Big sagebrush, Gardner’s saltbush, and birdfoot sagebrush are conspicuous elements of this state, and make up 20% of the annual production. Big sagebrush may become dominant on some areas with absence of fire. A variety of forbs also occurs in this state and plant diversity is high (see Plant Composition Table).
The total annual production (air-dry weight) of this state is about 300 pounds per acre, but it can range from about 150 lbs. /acre in unfavorable years to about 375 lbs. /acre in above average years.
The state is extremely stable and well adapted to the Northern Intermountain Desertic Basins climatic conditions. The diversity in plant species allows for high drought resistance. This is a sustainable plant community (site/soil stability, watershed function, and biologic integrity
Transitions or pathways leading to other plant communities are as follows:
• Moderate, continuous season-long grazing will convert the plant community to the Perennial Grass/Mixed Sagebrush Plant Community. Prolonged drought will exacerbate this transition.
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State 2
Perennial Grass/Mixed Sagebrush Plant Community
Community 2.1
Perennial Grass/Mixed Sagebrush Plant Community
Historically, this plant community evolved under grazing and a low fire frequency. Currently, it is found under moderate, season-long grazing by livestock and will be exacerbated by prolonged drought conditions. In addition, the fire regime for this site has been modified and extended periods without fire is now common. This plant community is still dominated by cool-season grasses, while short warm-season grasses and miscellaneous forbs account for the balance of the understory.
A variety of shrubs is now a conspicuous part of the overall production.
Dominant grasses include western wheatgrass, and bottlebrush squirreltail. Grasses and grass-like species of secondary importance include blue grama, Sandberg bluegrass and threadleaf sedge. Forbs commonly found in this plant community include scarlet globemallow, wild onion, smooth woodyaster, leafy wildparsley, and Hood’s phlox. Big sagebrush, birdfoot sagebrush and Gardner’s saltbush dominate the overstory. Big sagebrush and birdfoot sagebrush can make up to 20% of the annual production. Plains pricklypear cactus can also occur.
When compared to the Historic Climax Plant Community, big sagebrush and blue grama have increased. Plains pricklypear cactus will also have increased, but occurs only in small patches. Indian ricegrass has decreased and may occur in only trace amounts under the sagebrush canopy or within the patches of pricklypear. In addition, winterfat may or may not have changed depending on the season of use.
The total annual production (air-dry weight) of this state is about 200 pounds per acre, but it can range from about 100 lbs. /acre in unfavorable years to about 300 lbs. /acre in above average years.
This plant community is resistant to change. The herbaceous species present are well adapted to grazing; however, species composition can be altered through long-term overgrazing. The herbaceous component is mostly intact and plant vigor and replacement capabilities are sufficient. Water flow patterns and litter movement may be occurring but only on steeper slopes. Incidence of pedestalling is minimal. Soils are mostly stable and the surface shows minimum soil loss. The watershed is functioning and the biotic community is intact.
Transitions or pathways leading to other plant communities are as follows:
• Prescribed grazing or possibly long-term prescribed grazing, will convert this plant community to the HCPC. The probability of this occurring is high especially if rotational grazing along with short deferred grazing is implemented as part of prescribed method of use. In addition, the removal of fire suppression will allow a somewhat natural fire regime to reoccur to more easily transition between this plant community and the HCPC. A prescribed fire treatment can be useful to hasten this transition if desired.
• Frequent and severe grazing on more saline soils will convert the plant community to the Salt Tolerant Shrub/Bare Ground Plant Community.
• Frequent and severe grazing (yearlong grazing) on less saline soils, will convert the plant community to the Mixed Sagebrush/Blue Grama Sod Plant Community.
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State 3
Mixed Sagebrush/Blue Grama Sod Plant Community
Community 3.1
Mixed Sagebrush/Blue Grama Sod Plant Community
This plant community is the result of frequent and severe yearlong grazing. Soils on these sites are usually less saline. It is dominated by a dense sod of blue grama and includes a mosaic shrub overstory. Big sagebrush may be present but usually birdfoot sagebrush is the most important shrub in this plant community. Pricklypear cactus can become dense in areas so that livestock cannot graze forage growing within the cactus clumps.
When the historic climax plant community is replaced by warm season grasses total annual production is reduced and the ability of perennial cool season grasses is not able to remain as part of the plant composition.
The total annual production (air-dry weight) of this state is about 75 pounds per acre, but it can range from about 50 lbs. /acre in unfavorable years to about 150 lbs. /acre in above average years.
This state is relatively stable and protected from excessive erosion where the sod cover is excessive. The sod formed by these grasses is resistant to water infiltration. While the soil is protected by this sod, excessive runoff may occur off-site and on-site where sod is patchier. As a result, rills or other more severe erosion can occur on unprotected areas. The watershed may or may not be functioning, as runoff may affect adjoining sites. The biotic integrity of this plant community is not intact. Plant diversity is extremely low.
Transitional pathways leading to other plant communities are as follows:
• Grazing land mechanical treatment (chiseling, etc.) followed by prescribed grazing and, if necessary, seeding, will return this plant community to near Historic Climax Plant Community.
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State 4
Salt Tolerant Shrub/Bare Ground Plant Community
Community 4.1
Salt Tolerant Shrub/Bare Ground Plant Community
This plant community can occur on sites subjected to continuous yearlong grazing and where soils are influenced by elevated amounts of soluble salts. Salt tolerant shrubs are a significant component of the plant community and the preferred cool season grasses have been eliminated or greatly reduced. Wyoming big sagebrush makes up a minor component of the plant community.
This site is dominated by an overstory of salt tolerant shrubs, such as greasewood, birdfoot sagebrush and saltbushes, but can vary widely in their composition and production. This variation results from the varying quantity of soluble salts present in the soils and the availability of shrubs to occupy the site. Big sagebrush and rubber rabbitbrush are present but are mostly in small patches.
Perennial cool season mid-grasses have been removed leaving mostly patches of blue grama and annuals. Cheatgrass and weedy annual forbs such as halogeton, Russian thistle, and kochia, will occupy the site if a seed source is available. Noxious weeds such as Russian knapweed may also invade this state. Plant diversity is moderate to poor. When compared to the HCPC, grass production has diminished but is compensated by the increase in shrub production.
The interspaces between plants have expanded leaving the amount of bare ground more prevalent. Surface salts have increased, especially on sites dominated by greasewood and saltbushes. The leaves of these plants contain high amounts of sodium and other salts, and when shed these soluble salts are transferred to the soils underneath the plants. Consequently, the soil can exhibit wide variations in soluble salts, which can explain the variation in shrub composition.
The total annual production (air-dry weight) of this state is about 150 pounds per acre, but it can range from about 75 lbs. /acre in unfavorable years to about 200 lbs. /acre in above average years.
This plant community is resistant to change. These areas are actually more resistant to fire as less fine fuels are available and the bare ground between the shrubs has increased. Continued frequent and severe grazing or the removal of grazing does not seem to affect the composition or structure of the plant community. Plant diversity is moderate to poor. The biotic integrity of this state is mostly dysfunctional because of the predominant salt tolerant shrub overstory and absence of perennial cool season grasses.
Soil erosion is accelerated because of increased bare ground. Water flow patterns and pedestalling are obvious. Infiltration is reduced and runoff is increased. Rill channels may be noticeable in the interspaces and gullies may be establishing where rills have concentrated down slope.
Transitions or pathways leading to other plant communities are as follows:
• Prescribed grazing or possibly long-term prescribed grazing, will convert this plant community to the Salt Tolerant Shrub/Rhizomatous Wheatgrass Plant Community. Recovery to near Historic Climax Plant Community condition is difficult to impossible due to the resistance of these shrubs to herbicides and other brush management techniques. In addition, the increase in surface salts has had accumulated effects on the soil so most of the herbaceous plants associated with the HCPC are no longer suitable for this site. The most notable exception is the rhizomatous wheatgrasses and bottlebrush squirreltail. Soil remediation to reduce the surface salts is not recommended, as this is mostly ineffective and extremely costly. Seeding more salt-tolerant native grasses and forbs will improve the productivity of site and plant cover.
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State 5
Salt Tolerant Shrub/Rhizomatous Wheatgrasses Plant Community
Community 5.1
Salt Tolerant Shrub/Rhizomatous Wheatgrasses Plant Community
This plant community can occur where the Salt Tolerant Shrub/Bare Ground Plant Community is rested and a prescribed grazing management practice is implemented. Salt tolerant shrubs remain a significant component of the plant community, but preferred cool season grasses have reestablished.
This site is dominated by an overstory of salt tolerant shrubs, such as birdfoot sagebrush, saltbushes, and greasewood, but can exhibit a wide variety of shrub composition and production. Some perennial cool season mid-grasses have once again reestablished such as rhizomatous wheatgrasses and bottlebrush squirreltail. Other important grasses include Sandberg bluegrass and blue grama. Patches of annuals such as cheatgrass and other weedy annual forbs such as halogeton, Russian thistle, and kochia, will persist on this site. Noxious weeds such as Russian knapweed may also remain if not treated. The interspaces between plants will have diminished in size.
The total annual production (air-dry weight) of this state is about 200 pounds per acre, but it can range from about 125 lbs. /acre in unfavorable years to about 300 lbs. /acre in above average years.
This plant community is mostly resistant to change, but species composition can be altered through long-term overgrazing. The herbaceous component is stable and plant vigor and replacement capabilities are sufficient. The watershed may or may not be functioning and the biotic community is not intact because of the predominant salt tolerant shrub overstory. Plant diversity is moderate
Soils are mostly stable and recent soil loss is minimal. This should not be confused with evidence of remnant erosion. Water flow patterns and litter movement is stable but is still occurring on steeper slopes. Incidence of pedestalling is improving.
Transitions or pathways leading to other plant communities are as follows:
• Frequent and severe grazing will convert the plant community to the Salt Tolerant Shrub/Bare Ground Plant Community.
• Recovery to near Historic Climax Plant Community condition is difficult to impossible due to the resistance of these shrubs to herbicides and other brush management techniques. In addition, the increase in surface salts has had accumulated effects on the soil so most of the herbaceous plants associated with the HCPC are no longer suitable for this site. The most notable exception is the rhizomatous wheatgrasses and bottlebrush squirreltail. Soil remediation to reduce the surface salts is not recommended, as this is mostly ineffective and extremely costly. Seeding more salt-tolerant grasses and forbs will improve the productivity and plant cover of the site, but will not improve the biotic integrity.
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