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Ecological site F090BY008WI

Moist Sandy Bedrock Upland

Last updated: 11/16/2023
Accessed: 11/23/2024

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General information

Provisional. A provisional ecological site description has undergone quality control and quality assurance review. It contains a working state and transition model and enough information to identify the ecological site.

MLRA notes

Major Land Resource Area (MLRA): 090B–Central Wisconsin Thin Loess Dissected Till Plain

The Wisconsin and Minnesota Thin Loess MLRA, Northern and Southern Parts (90A and 90B) correspond closely to the North Central Forest and the Forest Transition Ecological Landscapes, respectively. Some of the following brief overview is borrowed from the Wisconsin Department of Natural Resources ecological landscape publications (2015).

The Wisconsin and Minnesota Thin Loess MLRA, Northern and Southern Parts (90A and 90B) is an extensive glacial landscape that comprised of over 11.1 million acres (17,370 sq mi) throughout central and northern Wisconsin – about 27% of the total land area in the state. This glacial landscape is comprised of a heterogenous mix of loess-capped ground moraines, end moraines with eskers and ice-walled lake plains, and pitted, unpitted, and collapsed outwash plains sometimes interspersed with drumlins from the Illinoian and Pre-Illinoian glaciations. The entire area has been glaciated and nearly all of it is underlain by dense glacial till that impedes drainage. An extensive morainal system – the Perkinstown end moraine – spans most of the width of northern Wisconsin and divides the Northern and Southern Parts of this large landscape. This moraine, which has been sliced by outwash in many places, marks the southernmost extent of the Wisconsin glaciation (Wisconsin’s most recent glacial advance).

North of the Perkinstown morainal system is a loess plain, with a loess mantle 6 to 24 inches thick. The northernmost edge of this landscape is an undulating till and outwash plain with materials deposited by the Chippewa Lobe. Drumlins are common in the northern and northeastern portions. The drumlins are oriented towards the southwest and formed during a glacial episode prior to the most recent glacial advance. Some are covered with glacial till. Pitted, unpitted, and collapsed outwash plains fill the spaces between drumlins. Detached from the major land mass to the northeast is the hummocky Hayward collapsed end moraines, where swamps, ice-walled lake plains, and eskers are common.

Most of the MLRA to the south of the Perkinstown morainal system is an extensive ground moraine with some proglacial stream features including pitted outwash plains, terraces, and fans. A layer of loess 6 to 47 inches thick covers much of the area. Like the Northern Part, all areas of the Southern Part of this MLRA were glaciated, although the southcentral portion is a relatively older till plain with materials from the Illinoian and pre-Illinoian glaciations, not the most recent Wisconsin glaciation. The landforms in the southcentral portion are highly variable. Much of the area topography is controlled by underlying bedrock. Sandstone outcrops and pediments can be found here. Some of the most southern portions of the MLRA are mixed glacial deposits and residuum.

The land surface of the southeastern portion was formed by many small glacial advances and retreats. Morainal ridges protrude through an erosional, pitted outwash-mantled surface. These parallel ridges run in a northeast to southwest orientation and are dissected by many steams.

The continental climate of this MLRA is typical of northcentral Wisconsin, with cold winters and warm summers. The southern boundary of this MLRA straddles Wisconsin’s Tension Zone, a zone of transition between Wisconsin’s northern and southern ecological landscapes. Historically, the mesic forests were dominated by eastern hemlock (Tsuga canadensis), sugar maple (Acer saccharum), and yellow birch (Betula alleghaniensis).

Classification relationships

Major Land Resource Area (MLRA): Wisconsin and Minnesota Thin Loess and Till (Northern and Southern Parts - 90A and 90B)
USFS Subregions: Rib Mountain Rolling Ridges (212Qd), Lincoln Formation Till Plain - Hemlock Hardwoods (212Qc), Lincoln Formation Till Plain - Mixed Hardwoods (212Qb), Rosemont Baldwin Plains and Moraines (222Md)
Wisconsin DNR Ecological Landscapes: Forest Transition, Western Prairie

Ecological site concept

The Moist Sandy Bedrock Upland ecological site is located on ground moraines, hills, and rock pediments. It’s found primarily in the southeast corner of MLRAs 90A and 90B where bedrock is shallower. These sites are characterized by moderately deep to deep, somewhat poorly drained soils that formed in a variety of parent materials including alluvium, till, and residuum. Soils are underlain with bedrock including shale, mica schist, interbedded sandstone and shale, greenstone and/or granite, and igneous and metamorphic rock. Precipitation, runoff from adjacent uplands, and groundwater discharge are the primary sources of groundwater. Soils range from extremely acid to neutral.

Moist Sandy Bedrock Upland is differentiated from other ecological sites based on drainage and moderately deep profile. Other somewhat poorly drained sands have soils that are greater than 80 inches in depth. The bedrock both perches the water table and restricts root growth. These sites are more vulnerable to tree tips. Other somewhat poorly drained sites have loamy or clayey deposits. Sands have lower pH and available water capacity than loamy and clayey sites, which can limit vegetative growth. The somewhat poor drainage sets this site apart from other sandy sites.

Associated sites

F090BY007WI	Wet Clayey Lowland Wet Clayey Lowland form in deep, loamy to clayey deposits derived from a mixture of alluvium, residuum, till, or lacustrine sources. These sites have a seasonally high water table at the surface, and some are subject to occasional ponding. Sustained saturation is enough for hydric conditions to occur. They are wetter and occur lower on the drainage sequence than Moist Sandy Bedrock Upland.
F090BY014WI	Loamy Bedrock Upland Loamy Bedrock Upland consist of loamy till, alluvium, or eolian deposits underlain by sandy to loamy residuum. Some sites may also contain sandy outwash or clayey pedisediment. Bedrock contact occurs within two meters of the surface. They have a seasonally high water table within one meter of the surface, though they don't remain saturated for extended periods of time. They are drier and occur higher on the drainage sequence than Moist Sandy Bedrock Upland.
F090BY021WI	Dry Loamy Upland Dry Loamy Upland consist of deep sandy to loamy outwash, alluvium, or till. The water table is deeper than two meters year-round. They are drier and occur higher on the drainage sequence than Moist Sandy Bedrock Upland.

F090BY007WI

Wet Clayey Lowland

Wet Clayey Lowland form in deep, loamy to clayey deposits derived from a mixture of alluvium, residuum, till, or lacustrine sources. These sites have a seasonally high water table at the surface, and some are subject to occasional ponding. Sustained saturation is enough for hydric conditions to occur. They are wetter and occur lower on the drainage sequence than Moist Sandy Bedrock Upland.

F090BY014WI

Loamy Bedrock Upland

Loamy Bedrock Upland consist of loamy till, alluvium, or eolian deposits underlain by sandy to loamy residuum. Some sites may also contain sandy outwash or clayey pedisediment. Bedrock contact occurs within two meters of the surface. They have a seasonally high water table within one meter of the surface, though they don't remain saturated for extended periods of time. They are drier and occur higher on the drainage sequence than Moist Sandy Bedrock Upland.

F090BY021WI

Dry Loamy Upland

Dry Loamy Upland consist of deep sandy to loamy outwash, alluvium, or till. The water table is deeper than two meters year-round. They are drier and occur higher on the drainage sequence than Moist Sandy Bedrock Upland.

Similar sites

F090BY011WI	Moist Loamy Lowland Moist Loamy Lowland consist of deep sandy and loamy deposits derived from a mixture of alluvium, residuum, till, or lacustrine sources. The finer textures allow the soil to stay moist - but not saturated - for sustained periods during the growing season. They share their particle size and drainage class with Moist Sandy Bedrock Upland. The vegetative communities they support are very similar.
F090BY010WI	Moist Loamy Lowland with Carbonates Moist Loamy Lowland With Carbonates consists of deep loamy till, sometimes with a loess mantle. Carbonates are present in these soils. The finer textures allow the soil to stay moist - but not saturated - for sustained periods during the growing season. These sites share their particle size and drainage class.
F090BY012WI	Moist Clayey Lowland Moist Clayey Lowland sites consist of deep clayey lacustrine deposits. The finer textures perch the water table. These soils remain moist - but not saturated - throughout much of the growing season. They share landscape position and drainage class with Moist Sandy Bedrock Upland, sometime with similar textures. The vegetative communities they support are very similar.
F090BY004WI	Loamy Floodplain Loamy Floodplain are found exclusively on floodplains in loamy alluvium, sometimes underlain by sandy alluvium. Soils are very poorly to moderately well drained and are subject to flooding. Some sites may be saturated for long enough for hydric conditions to occur. They share their particle size and sometimes their drainage class with Moist Sandy Bedrock Upland. The vegetative communities they support are very similar.

Table 1. Dominant plant species

Tree	(1) Acer saccharum (2) Tsuga canadensis
Shrub	(1) Corylus cornuta
Herbaceous	(1) Arisaema (2) Maianthemum canadense

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Physiographic features

These sites formed on ground moraines, hills, and rock pediments. Slopes range from 0 to 8 percent. Sites are in the footslope position.

These sites are not subject to ponding or flooding. Some soils have an apparent seasonally high water table (endosaturation) at a depth of 6 to 36 inches. Other sites have a perched seasonally high water table (episaturation) at a depth of 6 to 18 inches. The water table may drop below 80 inches during dry conditions on all sites. Runoff is negligible to very high.

Table 2. Representative physiographic features

Hillslope profile	(1) Footslope
Slope shape across	(1) Concave
Slope shape up-down	(1) Linear
Landforms	(1) Ground moraine (2) Hill (3) Rock pediment
Runoff class	Low to very high
Flooding frequency	None
Ponding frequency	None
Elevation	180 – 275 m
Slope	0 – 8%
Water table depth	15 – 91 cm
Aspect	Aspect is not a significant factor

Climatic features

The climate of the expansive Wisconsin and Minnesota Thin Loess and Till Plain is highly variable. The eco-climatic zone (the “Tension Zone”) that runs southeast-northwest across the state splits the MLRA. In general, the MLRA has cold winters and warm summers with an adequate amount of precipitation. Near Lake Superior, precipitation and temperature tend to increase. The far western section of the MLRA, known as the western prairie ecological landscape by the Wisconsin DNR, has warmer temperatures compared to the rest of the MLRA because it falls below the eco-climatic zone. The soil moisture regime of MLRA is udic (humid climate). The soil temperature regime is frigid and cryic.

The average annual precipitation for this ecological site is 31 inches. The average annual snowfall is 50 inches. The annual average maximum and minimum temperatures are 53°F and 33°F, respectively.

Table 3. Representative climatic features

Frost-free period (characteristic range)	95-116 days
Freeze-free period (characteristic range)	129-140 days
Precipitation total (characteristic range)	787-813 mm
Frost-free period (actual range)	93-117 days
Freeze-free period (actual range)	119-147 days
Precipitation total (actual range)	686-838 mm
Frost-free period (average)	104 days
Freeze-free period (average)	134 days
Precipitation total (average)	787 mm

Characteristic range

Actual range

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Figure 1. Monthly precipitation range

Characteristic range

Actual range

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Figure 2. Monthly minimum temperature range

Characteristic range

Actual range

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Figure 3. Monthly maximum temperature range

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Figure 4. Monthly average minimum and maximum temperature

Figure 5. Annual precipitation pattern

Figure 6. Annual average temperature pattern

Climate stations used

(1) WINTER [USC00479304], Ojibwa, WI
(2) LUCK [USC00474894], Luck, WI
(3) HOLCOMBE [USC00473698], Holcombe, WI
(4) LAONA 6 SW [USC00474582], Laona, WI
(5) SANDSTONE 6 W [USW00054932], Hinckley, MN
(6) ISLE 12N [USC00214103], Isle, MN

Influencing water features

Water is received through precipitation, runoff from adjacent uplands, and groundwater discharge. Water levels are greatly influenced by precipitation rates and runoff from upland sites. Water leaves the site primarily through runoff, evapotranspiration, and groundwater recharge. Subsurface outflow may occur when water perches on bedrock.

Wetland description

Permeability of the soils is impermeable to moderately slow.
Hydrologic Group: D, A/D, B/D, C/D
Hydrogeomorphic Wetland Classification: None
Cowardin Wetland Classification: None

Soil features

These sites are represented by the Derinda Variant, Dolph, Kert, Magroc, Merrillan, Mora, Rietbrock, and Shullsburg soil series. Derinda Variant is classified as an Aquollic Hapludalf; Dolph is an Aeric Glossaqualf; Kert, Magroc, Mora, and Rietbrock are Aquic Glossaqualfs; Merrillan is an Ultic Epiaquod; Shullsburg is an Aquic Argiudoll.

These soils formed in various parent materials including loess, sandy, loamy, and clayey residuum, sandy or silty alluvium, and loamy till. Soils are moderately deep to very deep with bedrock contact occurring between 66 and 165 cm. Sites are somewhat poorly drained.

The surface of these sites is silt loam or moderately decomposed plant material. Subsurface horizon textures include silt loam, sandy loam, fine sandy loam, clay loam, sandy clay loam, silty clay loam, loam, silty clay, clay, loamy sand, and sand. Soil pH ranges from extremely acid to neutral with values of 3.60 to 7.20. Surface fragments less than 3 inches may be present up to 7 percent cover, and fragments greater than 3 inches may be present up to 3 percent cover. Subsurface fragments less than 3 inches may be present up to 21 percent volume, and fragments greater than 3 inches may be present up to 18 percent. Carbonates may be present up to 10 percent beginning at 74 cm.

Table 4. Representative soil features

Parent material	(1) Alluvium (2) Eolian deposits (3) Till (4) Sandstone and shale (5) Igneous and metamorphic rock
Surface texture	(1) Silt loam
Drainage class	Somewhat poorly drained
Permeability class	Very slow to moderately slow
Soil depth	66 – 165 cm
Surface fragment cover <=3"	0 – 7%
Surface fragment cover >3"	0 – 5%
Available water capacity (0-154.9cm)	3.1 – 8.92 cm
Calcium carbonate equivalent (0-100.1cm)	0 – 10%
Soil reaction (1:1 water) (0-100.1cm)	3.6 – 7.2
Subsurface fragment volume <=3" (Depth not specified)	0 – 21%
Subsurface fragment volume >3" (Depth not specified)	0 – 18%

Ecological dynamics

Historically, mature forests on this ecological site were dominated by shade tolerant sugar maple and hemlock, often with an admixture of yellow birch (Wilde, 1933, Finley, 1976). This association was self-maintained with new cohorts of advance regeneration gaining canopy status through gaps formed by small-scale disturbances and natural mortality in the dominant canopy. Scattered large individuals of less shade tolerant white pine also were common component of mesic hardwood forests. These presumably became established following relatively rare disturbances that included fire (Schulte and Mladenoff, 2005).

Current stands on this Ecological Site represent the entire array of potential successional stages from pure aspen, or aspen-white birch, stands to sugar maple dominated mixed northern hardwoods stands. Succession to sugar maple dominance is evident everywhere that seed sources are present. However, hemlock regeneration is scarce. In old forests, hemlock finds optimal conditions for germination and seedling establishment on rotten logs, stumps and mounds that normally have warmer surfaces and better moisture retention than the forest floor (USDA, 1990). Most present forest communities lack these conditions.

State and transition model

More interactive model formats are also available. View Interactive Models
Click on state and transition labels to scroll to the respective text

Ecosystem states

1. Reference State

2. Deciduous Forest State

3. Agricultural State

T1A	-	Stand replacing disturbance that includes fire.
T1B	-	Removal of forest cover and tilling for agricultural crop production.
R2	-	Deciduous forest community is slowly invaded by conifers.
T2A	-	Removal of forest cover and tilling for agricultural crop production.
R3A	-	Cessation of agricultural practices leads to natural reforestation, or site is replanted.
T3A	-	Cessation of agricultural practices leads to natural reforestation, or site is replanted.

State 1 submodel, plant communities

1.1. Advanced Success Phase

1.2. Rejuvenated Phase

1.1A	-	Light to moderate intensity fires, blow-downs, ice storms.
1.2A	-	Disturbance-free period for 30+ years.

State 2 submodel, plant communities

2.1. Deciduous Forest Phase

State 3 submodel, plant communities

3.1. Agricultural Phase

State 1
Reference State

The reference plant community is categorized as mesic forest community dominated by mixed deciduous species, primarily sugar maple (Acer saccharum), and sporadic occurrence of several conifer species. Although forest communities can vary greatly in terms of species composition and stand structure, depending on type, degree, and frequency of disturbance, two common phases predominate:

Community 1.1
Advanced Success Phase

In the absence of major, stand-replacing disturbance this community is dominated by sugar maple, yellow birch (Betula alleghaniensis) and eastern hemlock (Tsuga canadensis. This was the most common condition in pre-European settlement forests. The tree sapling and shrub layer in this community is not well developed due to dense shade created by multi-story tree canopy. Sugar maple saplings dominate the shrub layer, but other shrubs including beaked hazelnut (Corylus cornuta) and cherries (Prunus, spp.) are common with low coverage. The herb layer is relatively species rich, but moderate in abundance. The dominant herbs typically include Jack-in-the-pulpit (Arisaema triphyllum) and Canada mayflower (Maianthemum canadense). Other species include Virginia creeper (Parthenocissus quinquefolia) and hairy Solomon’s seal (Polygonatum pubescens). It is important to note that in most current mature stands, hemlock is significantly under-represented compared to historic conditions. Apparently, this lack of hemlocks is due to seed source elimination during the early logging era and herbivory by currently high white tail deer populations.

Dominant plant species

sugar maple (Acer saccharum), tree
eastern hemlock (Tsuga canadensis), tree
yellow birch (Betula alleghaniensis), tree
beaked hazelnut (Corylus cornuta), shrub
Jack in the pulpit (Arisaema), other herbaceous
Canada mayflower (Maianthemum canadense), other herbaceous

Community 1.2
Rejuvenated Phase

Disturbances described in Pathway 1.1A lead to increased species and structural diversity of the forest community. Depending on seed source, red oak and red maple regenerate in the canopy openings and in time join sugar maple and hemlock in the dominant canopy. The relative density of the shrub and herb layers also increases during this stage. Species composition remains relatively unchanged, but abundance changes can be significant. Particularly beaked hazelnut can form dense thickets. Many other herb species that were present with very low abundance in the advanced-succession community typically form much larger population clusters.

Dominant plant species

sugar maple (Acer saccharum), tree
red maple (Acer rubrum), tree
northern red oak (Quercus rubra), tree
beaked hazelnut (Corylus cornuta), shrub
Jack in the pulpit (Arisaema), other herbaceous
Canada mayflower (Maianthemum canadense), other herbaceous

Pathway 1.1A
Community 1.1 to 1.2

Natural mortality in the oldest age classes—sporadic small-scale blow-downs and ice storms—create openings for entry of mid-tolerant species such as red oak and red maple.

Pathway 1.2A
Community 1.2 to 1.1

In the absence of canopy reducing disturbances natural succession leads to community dominance by the most shade-tolerant species resulting in return to community phase 1.1.

State 2
Deciduous Forest State

Post disturbance pioneer community of aspen and paper birch with mixtures of other species from available seed sources.

Community 2.1
Deciduous Forest Phase

Pure, or mixed, aspen – paper birch community replaces the reference state community. If seed source is present, red maple and red oak readily becomes member of this community.

Dominant plant species

quaking aspen (Populus tremuloides), tree
red maple (Acer rubrum), tree
northern red oak (Quercus rubra), tree
beaked hazelnut (Corylus cornuta), shrub
Canada mayflower (Maianthemum canadense), other herbaceous
goldenrod (Oligoneuron), other herbaceous

State 3
Agricultural State

Indefinite period of applying agricultural practices.

Community 3.1
Agricultural Phase

Indefinite period of applying agricultural practices. Crops likely include alfalfa, corn, soybeans, and hay or pasture. It is possible that some areas are or have been in ginseng production as well.

Transition T1A
State 1 to 2

Major stand-replacing disturbance. In pre-European settlement time, the event was most often a severe blow down, sometimes followed by fires. Such blow downs have been estimated to occur in this part of Wisconsin every 300 to 400 years (Schulte and Mladenoff, 2005). Blow downs may occur more frequently on these sites because of the shallow bedrock. In post settlement virtually every acre has been logged either by clear cutting or successive cuts targeting species marketable at that time. Post logging slash fires also have been a significant factor in most areas. These disturbances created the environment suitable for natural regeneration of many shade-intolerant species and for commercial planting.

Transition T1B
State 1 to 3

Removal of forest cover, tilling and application of other agricultural techniques to grow agricultural crops.

Restoration pathway R2
State 2 to 1

Deciduous forest community is slowly invaded by conifers.

Transition T2A
State 2 to 3

Removal of forest cover, tilling and application of other agricultural techniques to grow agricultural crops.

Restoration pathway R3A
State 3 to 1

Abandonment of agricultural practices and allowing natural vegetation to colonize the site or apply artificial afforestation. The time required for forest community to reach the reference state conditions may exceed 100 years.

Transition T3A
State 3 to 2

Abandonment of agricultural practices and allowing natural vegetation to colonize the site or apply artificial afforestation.

Additional community tables

Supporting information

Inventory data references

Plot and other supporting inventory data for site identification and community phases is located on a NRCS North Central Region shared and one drive folder. University Wisconsin-Stevens Point described soils, took photographs, and inventoried vegetation data at community phases within the reference state. The data sources include WI ESD Plot Data Collection Form - Tier 2, Releve Method, NASIS pedon description, NRCS SOI 036, photographs, and Kotar Habitat Types.
Habitat Types of N. Wisconsin (Kotar, 2002): The sites of this ES keyed out to three habitat types: Acer/Hydrophyllum-Impatiens (AHI); Acer-Tsuga/Athyrium-Onoclea (ATAtOn); Acer-Tsuga/Maianthemum (ATM)
Biophysical Settings (Landfire, 2014): This ES is largely mapped as Laurentian-Acadian Northern Hardwoods Forest, Eastern Cool Temperate Row Crop, and Eastern Cool Temperate Close Grown Crop
WDNR Natural Communities (WDNR, 2015):

Other references

Cleland, D.T.; Avers, P.E.; McNab, W.H.; Jensen, M.E.; Bailey, R.G., King, T.; Russell, W.E. 1997. National Hierarchical Framework of Ecological Units. Published in, Boyce, M. S.; Haney, A., ed. 1997. Ecosystem Management Applications for Sustainable Forest and Wildlife Resources. Yale University Press, New Haven, CT. pp. 181-200.

County Soil Surveys from St. Croix, Polk, Barron, Rusk, Chippewa, Clark, Marathon, Taylor, Price, Sawyer, Burnett, Washburn, Douglas, Bayfield, Ashland, Lincoln, Oneida, Langlade, Shawano, Menominee, Forest, Florence, Marinette, and Pierce Counties.

Curtis, J.T. 1959. Vegetation of Wisconsin: an ordination of plant communities. University of Wisconsin Press, Madison. 657 pp.

Davis, R.B. 2016. Bogs and Fens, A Guide to the Peatland Plants of Northeastern United States and Adjacent Canada. University Press of New England, Hanover and London. 296 pp.

Finley, R. 1976. Original vegetation of Wisconsin. Map compiled from U.S. General Land Office notes. U.S. Forest Service, North Central Forest Experiment Station, St. Paul, Minnesota.

Hvizdak, David. Personal knowledge and field experience.

Jahnke, J. and Gienccke, A. 2002. MLRA 92 Clay Till Field Investigations. Summary of field day investigations by Region 10 Soil Data Quality Specialists.

Kotar, J. 1986. Soil – Habitat Type relationships in Michigan and Wisconsin. J. For. and Water Cons. 41(5): 348-350.

Kotar, J., J.A. Kovach and G. Brand. 1999. Analysis of the 1996 Wisconsin Forest Statistics by Habitat Type. U.S.D.A. For. Serv. N.C. Res. Stn. Gen. Tech. Rept. NC-207.

Kotar, J., J. A. Kovach, and T. L. Burger. 2002. A Guide to Forest Communities and Habitat Types of Northern Wisconsin. Second edition. University of Wisconsin-Madison, Department of Forest Ecology and Management, Madison.

Kotar, J., and T. L. Burger. 2017. Wetland Forest Habitat Type Classification System for Northern Wisconsin: A Guide for Land Managers and landowners. Wisconsin Department of Natural Resources, PUB-FR-627 2017, Madison.

Martin, L. 1965. The physical geography of Wisconsin. Third edition. The University of Wisconsin Press, Madison.

McNab, W.H. and P.W. Avers. 1994. Ecological Subregions of the United States: Section Descriptions. USDA For. Serv. Pun. WO-WSA-5, Washington, D.C.

NatureServe. 2018. International Ecological Classification Satandard: Terrestrial Ecological Classifications. NautreServe Centreal Databases. Arlington, VA. U.S.A. Data current as of 28 August 2018.

Radeloff, V.C., D.J. Mladenoff, H.S. He and M.S. Boyce. 1999. Forest landscape change in Northwestern Wisconsin Pine Barrens from pre-European settlement to the present. Can. J. For. Res. 29: 1649-1659.

Schulte, L.A., and D.J. Mladenoff. 2001. The original U.S. public land survey records: their use and limitations in reconstructing pre-European settlement vegetation. Journal of Forestry 99:5–10.

Schulte, L.A., and D.J. Mladenoff. 2005. Severe wind and fire regimes in northern forests: historical variability at the regional scale. Ecology 86(2):431–445.

Soil Survey Staff. Input based on personal experience. Tim Miland, Scott Eversoll, Ryan Bevernitz, and Jason Nemecek.

Stearns, F. W. 1949. Ninety years change in a northern hardwood forest in Wisconsin. Ecology, 30: 350-58.

United States Department of Agriculture, Forest Service. 1989. Proceedings – Land Classification Based on Vegetation: Applications for Management. Gen. Tech. Report INT-527.

United States Department of Agriculture, Forest Service. 1990. Silvics of North America, Vol. 1, Hardwoods. Agricultural Handbook 654, Washington, D.C.

United States Department of Agriculture, Forest Service. 1990. Silvics of North America, Vol. 2, Conifers. Agricultural Handbook 654, Washington, D.C.

United States Department of Agriculture, Natural Resources Conservation Service. 2006. Land Resource and Major Land Resource Areas of the United Sates, the Caribbean, and the Pacific Basin. U.S. Department of Agriculture Handbook 296.

United States Department of Agriculture, Natural Resources Conservation Service. 2008. Hydrogeomorphic Wetland Classification System: An Overview and Modification to Better Meet the Needs of the Natural Resources Conservation Service. Technical Note No. 190-8-76. Washington D.C.

Wilde, S.A. 1933. The relation of soil and forest vegetation of the Lake States Region. Ecology 14: 94-105.

Wilde, S.A. 1976. Woodlands of Wisconsin. University of Wisconsin Cooperative Extension, Pub. G2780, 150 pp.

Wisconsin Department of Natural Resources. 2015. The ecological landscapes of Wisconsin: An assessment of ecological resources and a guide to planning sustainable management. Wisconsin Department of Natural Resources, PUB-SS-1131 2015, Madison.

Contributors

Bryant Scharenbroch, Assistant Professor at University of Wisconsin Stevens Point
Jacob Prater, Associate Professor at University of Wisconsin Stevens Point
John Kotar, Ecological Specialist, independent contractor

Approval

Suzanne Mayne-Kinney, 11/16/2023

Acknowledgments

NRCS contracted UWSP to write ecological sites in MLRA 90B, completed in 2021.

Rangeland health reference sheet

Interpreting Indicators of Rangeland Health is a qualitative assessment protocol used to determine ecosystem condition based on benchmark characteristics described in the Reference Sheet. A suite of 17 (or more) indicators are typically considered in an assessment. The ecological site(s) representative of an assessment location must be known prior to applying the protocol and must be verified based on soils and climate. Current plant community cannot be used to identify the ecological site.

Author(s)/participant(s)
Contact for lead author
Date	11/23/2024
Approved by	Suzanne Mayne-Kinney
Approval date
Composition (Indicators 10 and 12) based on	Annual Production