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Ecological site R092XY012WI

Moist Clayey Lowlands

Last updated: 4/09/2020
Accessed: 11/21/2024

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General information

Provisional. A provisional ecological site description has undergone quality control and quality assurance review. It contains a working state and transition model and enough information to identify the ecological site.

MLRA notes

Major Land Resource Area (MLRA): 092X–Superior Lake Plain

The Wisconsin portion of the Superior Lake Plain (MLRA 92) corresponds very closely to the Superior Coastal Plain Ecological Landscape published by Wisconsin Department of Natural Resources (WDNR 2015). The following brief overview of this MLRA is borrowed from that publication.

The Superior Coastal Plain is bordered on the north by Lake Superior and on the south by the Northwest Sands, Northwest Lowlands, and North Central Forest Ecological Landscapes. The total land area is approximately 1.2 million acres, which mostly consists of privately-owned forestland. The climate is strongly influenced by Lake Superior, resulting in cooler summers, warmer winters, and greater precipitation compared to more inland locations. The most extensive landform in this ecological landscape is a nearly level plain of lacustrine clays that slopes gently northward toward Lake Superior. The coastal plain is cut by deeply incised stream drainages and interrupted by the comparatively rugged Bayfield Peninsula.

During the Late Wisconsin glacial period, this area was covered with the advancing and retreating lobes of Superior and Chippewa. The landscape was rippled with moraines, but they were subdued by deposition of lacustrine materials. As the glaciers receded, glacial lakes riddled the landscape—most notably, Glacial Lake Duluth. The glacier receded eastward, exposing the western Lake Superior Basin. The ice covered the eastern basin, blocking the outlet of the lake, and continued to recede and contribute meltwaters that filled the glacial lake. The deep, red clays were deposited during this period of glacial lakes. The meltwaters from the glacier also contained sands which were deposited along the edge of the glacial lakes as beach deposits. Deep, narrow valleys have since been carved by rivers and streams flowing north into Lake Superior.

Historically, the Superior Coastal Plain was almost entirely forested. Various mixtures of eastern white pine (Pinus strobus), white spruce (Picea glauca), balsam fir (Abies balsamea), white birch (Betula papyrifera), balsam poplar (Populus balsamifera), quaking aspen (Populus tremuloides), and northern white-cedar (Thuja occidentalis) occurred on the fine-textured glacio-lacustrine deposits bordering much of the Lake Superior coast. Sandy soils, sometimes interlayered with clays, occur in some places. Such areas supported forests dominated by eastern white pine and red pine (Pinus resinosa). Eastern white pine was strongly dominant in some areas, according to mid-19th century notes left by surveyors of the federal General Land Office (Finley, R. 1976). Dry-mesic to wet-mesic northern hardwoods or hemlock-hardwood forests were prevalent on the glacial tills of the Bayfield Peninsula. Large peatlands occurred along the Lake Superior shoreline, associated with drowned river mouths.

Classification relationships

Habitat Types of N. Wisconsin (Kotar, 2002): One site in this ES keys out to Acer rubrum – Abies balsamea / Vaccinium spp. – Cornus canadensis [ArAbVCo], and one site keys out to Acer saccharum – Tsuga canadensis / Maianthemum canadense [ATM].

Biophysical Setting; Landfire (NatureServe, 2018): This is ES is mapped as Larentian – Acadian Northern Hardwoods Forest and Laurentian – Acadian Sub-boreal Mesic Balsam Fir-Spruce Forest. The ES is not well represented by any of these, but most similar to Sub-Boreal Mesic Balsam Fir - Spruce Forest.

WDNR Natural Communities (WDNR, 2015): This ES is not well represented by any of the described natural communities, but bears some resemblance to Northern Wet-Mesic Forest and Boreal Forest.

USFS Subregions: Superior-Ashland Clay Plain Subsection (212Ya); May contain small areas of Ewen Dissected Lake Plain Subsection (212Jo), Winegar Moraines Subsection (212Jc), Gogebic-Penokee Iron Range Subsection (212Jb), and NorthShore Highlands Subsection (212Lb)*

Major Land Resource Area (MLRA): Superior Lake Plain (92)

Ecological site concept

Moist Clayey Lowlands has a fairly wide extent throughout the MLRA, but most common in the Bayfield peninsula and the Apostle Islands. The sites are located on till plains and lake plains, with very fine and fine textures, sometimes with a sandy or loamy mantle over the clayey deposits. The fine materials cause a perched seasonally high water table that has a depth of 0 to 30 cm, but the water table can reach a depth of greater than 152 cm during dry periods. The primary source of water is precipitation, but runoff and groundwater also contribute. The soils range from strongly acid to strongly alkaline. Carbonates may be present beginning at 30 cm. The wide pH range is caused by soil differences—mores specifically, the soils with a sandy mantle are more acidic because of the coarse texture and lack of carbonates. This ES is wetter than Clayey Uplands, but not as wet as Wet Loamy or Clayey Lowlands. This ES differs from Moist Sandy Lowlands and Moist Loamy Lowlands based on its finer texture, clayey deposits, and its higher seasonally high water table. Typical vegetation includes balsam fir, red maple, sugar maple, paper birch, red oak, green ash, and quaking aspen. Ground flora includes ferns, beaked hazelnut, goldthread, American starflower, and Canada mayflower.

Associated sites

R092XY007WI	Wet Loamy or Clayey Lowlands These sites are poorly to very poorly drained soils that formed in mainly clayey deposits. Some sites may have a sandy or loamy mantle overlying a clayey deposit. Soils remain saturated throughout the year and frequently experience ponding and flooding in the spring and fall. Water table rarely drops below 30cm in drought conditions. Most sites have apparent carbonates, and pH ranges from strongly acid to moderately alkaline. HGM criteria: recharge, Depressional. These sites are often adjacent to Moist Clayey Lowlands but located on a lower landscape position in the drainage sequence.
R092XY015WI	Clayey Uplands These sites are deep, moderately well to well drained soils that formed in clayey till or glaciolacustrine deposits. Some sites have a sandy or loamy mantle. Sites have a seasonally high water table, but does not remain saturated for extended periods. Sites range from strongly acid to moderately alkaline, with carbonates present in many sites. These sites are often adjacent to Moist Clayey Lowlands but located on a higher landscape position in the drainage sequence.

R092XY007WI

Wet Loamy or Clayey Lowlands

These sites are poorly to very poorly drained soils that formed in mainly clayey deposits. Some sites may have a sandy or loamy mantle overlying a clayey deposit. Soils remain saturated throughout the year and frequently experience ponding and flooding in the spring and fall. Water table rarely drops below 30cm in drought conditions. Most sites have apparent carbonates, and pH ranges from strongly acid to moderately alkaline. HGM criteria: recharge, Depressional. These sites are often adjacent to Moist Clayey Lowlands but located on a lower landscape position in the drainage sequence.

R092XY015WI

Clayey Uplands

These sites are deep, moderately well to well drained soils that formed in clayey till or glaciolacustrine deposits. Some sites have a sandy or loamy mantle. Sites have a seasonally high water table, but does not remain saturated for extended periods. Sites range from strongly acid to moderately alkaline, with carbonates present in many sites. These sites are often adjacent to Moist Clayey Lowlands but located on a higher landscape position in the drainage sequence.

Similar sites

R092XY010WI	Moist Sandy Lowlands Moist Sandy Lowlands have a sandy mantle overlying finer glaciofluvial materials. The finer materials can cause episaturation in spring and fall, allowing the site to remain moist for some of the growing season, but does not remain saturated, nor does it have hydric conditions. These sites are found in a similar landscape to Moist Clayey Lowlands but are coarser textured and in a different drainage sequence.
R092XY011WI	Moist Loamy Lowlands These sites are somewhat poorly drained soils formed in various parent materials, but primarily are primarily loamy. The loamy texture causes the soil to remain moist for much of the growing season but does not remain saturated long enough to form hydric conditions. These sites are found in a similar landscape to Moist Clayey Lowlands but are coarser textured and in a different drainage sequence.

R092XY010WI

Moist Sandy Lowlands

Moist Sandy Lowlands have a sandy mantle overlying finer glaciofluvial materials. The finer materials can cause episaturation in spring and fall, allowing the site to remain moist for some of the growing season, but does not remain saturated, nor does it have hydric conditions. These sites are found in a similar landscape to Moist Clayey Lowlands but are coarser textured and in a different drainage sequence.

R092XY011WI

Moist Loamy Lowlands

These sites are somewhat poorly drained soils formed in various parent materials, but primarily are primarily loamy. The loamy texture causes the soil to remain moist for much of the growing season but does not remain saturated long enough to form hydric conditions. These sites are found in a similar landscape to Moist Clayey Lowlands but are coarser textured and in a different drainage sequence.

Table 1. Dominant plant species

Tree	(1) Acer rubrum (2) Abies balsamea
Shrub	(1) Corylus cornuta (2) Alnus incana ssp. rugosa
Herbaceous	(1) Pteridium aquilinum (2) Eurybia macrophylla

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Physiographic features

This site occurs on plains, drainageways, terraces, and footslopes located on till plains and lake plains. Landform shape is predominantly linear, but can be concave on footslopes. Elevation of the landforms range from 185 to 330 meters above sea level. Slopes are predominantly 0 to 6 percent, but will range up to 15 percent on footslopes. This site occurs on all slope aspects.

Table 2. Representative physiographic features

Landforms	(1) Lake plain > Terrace (2) Till plain > Drainageway
Runoff class	High to very high
Elevation	607 – 1,083 ft
Slope	15%
Water table depth	60 in
Aspect	Aspect is not a significant factor

Climatic features

The Moist Clayey Lowlands PESD is located throughout the MLRA, but most concentrated in the Bayfield Peninsula. The annual average precipitation ranges from 29-33 inches, with a range of 56-167 inches of annual average snowfall (PRISM, 1981-2010). The annual average minimum temperature ranges from 28-34oF, and the maximum temperature ranges from 46-52oF (PRISM, 1981-2010). The freeze free period ranges from 156-194 days, with an average of 171 days (Table 1). The frost-free period ranges from 130-166 days, with an average of 144 days (Table 1).

Table 3. Representative climatic features

Frost-free period (characteristic range)	80-113 days
Freeze-free period (characteristic range)	116-139 days
Precipitation total (characteristic range)	31-32 in
Frost-free period (actual range)	69-114 days
Freeze-free period (actual range)	107-142 days
Precipitation total (actual range)	31-33 in
Frost-free period (average)	98 days
Freeze-free period (average)	128 days
Precipitation total (average)	32 in

Characteristic range

Actual range

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Figure 1. Monthly precipitation range

Characteristic range

Actual range

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Figure 2. Monthly minimum temperature range

Characteristic range

Actual range

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Figure 3. Monthly maximum temperature range

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Figure 4. Monthly average minimum and maximum temperature

Figure 5. Annual precipitation pattern

Figure 6. Annual average temperature pattern

Climate stations used

(1) ASHLAND 3S [USC00470347], Ashland, WI
(2) ASHLAND EXP FARM [USC00470349], Ashland, WI
(3) BAYFIELD 6 N [USC00470603], Bayfield, WI
(4) MADELINE ISLAND [USC00474953], La Pointe, WI
(5) FOXBORO [USC00472889], Foxboro, WI
(6) PATTISON RANGER STAION [USC00476413], Foxboro, WI

Influencing water features

Water is received primarily through precipitation and from runoff from adjacent uplands. Water is lost from the site primarily through runoff and evapotranspiration.

Permeability of the soil ranges from impermeable to very slow. Runoff potential is high to very high. The hydrologic group of this site is primarily D, but where there is a sandy mantle over the clayey deposits the hydrologic group is C.

Enough water will percolate into some soil areas that will result in a perched seasonally high water table (episaturation) at a depth of 0 to 30 cm that may occur in any month, but will range to greater than 152 cm under dry conditions. Water that percolates into the soil is generally lost through plant uptake and evapotranspiration. Because of the permeability of the soil, there is very little to no ground water recharge.

Soil features

The soils of this site are represented by the Allendale, Badriver, Borea, Cuttre, Herbster, and Sedgwick soil series. These soils are predominantly classified as Glossaqualfs (Badriver, Borea, Cuttre, and Herbster), but includes sites that are Epiaquods (Allendale and Sedgwick).
This ecological site is characterized by very deep, somewhat poorly drained soils formed in clayey till or glaciolacustrine deposits (Badriver, Borea, Cuttre, and Herbster). Some areas include soil formed in a sandy glaciofluvial mantle over the clayey deposits (Allendale), while in other areas the soil formed in a loamy glaciofluvial mantle over the clayey deposits (Sedwick).
The average gravel content within the soil can be as much as 7 percent, while the average content of cobbles and stones can be as much as 1 percent. Soil reaction (pH) in the upper 100 cm ranges from strongly acid to strongly alkaline. Carbonates can be as shallow as 30 cm below the surface.

Table 4. Representative soil features

Parent material	(1) Till (2) Glaciolacustrine deposits
Surface texture	(1) Clay (2) Clay loam (3) Silt loam (4) Sandy loam (5) Loamy fine sand (6) Loamy sand
Drainage class	Somewhat poorly drained
Permeability class	Very slow
Available water capacity (0-60in)	3.47 – 5.17 in
Calcium carbonate equivalent (0-40in)	5 – 18%
Soil reaction (1:1 water) (0-40in)	5.3 – 8.5
Subsurface fragment volume <=3" (0-40in)	1 – 7%
Subsurface fragment volume >3" (0-40in)	1%

Ecological dynamics

Because of relatively poorly drained soils historic fire disturbance has likely been less frequent and less severe than on the better drained sites. This is evident by the presence (historic and current) of shade-tolerant and fire sensitive species such as red maple, balsam fir and white spruce. Aspen stands are common in current communities, but they are largely the result of fires associated with past logging. Red maple and balsam fir are the most obvious succeeding species, but white pine and white spruce may also become more important in the future as seed source availability increases. Although the shade-tolerant sugar maple occurs sporadically in some stands its competitive ability is reduced by excessive soil moisture and relatively low nutrient availability. For these reasons it is likely to remain only as a sporadic associate rather than the dominant component of mature forest communities as is typically the case on all mesic sites throughout northern Wisconsin.

State and transition model

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Click on state and transition labels to scroll to the respective text

Ecosystem states

1. Reference State

2. Deciduous Forest State

3. Agriculture State

T1A	-	Stand replacing disturbance that includes fire.
T1B	-	Removal of forest cover and tilling for agricultural crop production.
R2	-	Deciduous forest community is slowly invaded by conifers.
T2A	-	Removal of forest cover and tilling for agricultural crop production.
R3A	-	Cessation of agricultural practices leads to natural reforestation, or site is replanted.
R3B	-	Cessation of agricultural practices leads to natural reforestation, or site is replanted.

State 1 submodel, plant communities

1.1. Mature Community

1.2. Rejuvenated Community

1.1A	-	Blow-down, severe ice storm, or large-scale mortality in overstory.
1.2A	-	Advanced regeneration response to canopy disturbance.

State 2 submodel, plant communities

2.1. Aspen – Paper birch/Beaked hazelnut/Bracken fern-Big leaf aster

State 3 submodel, plant communities

3.1. Cultivated crops primarily hay

State 1
Reference State

Reference state is a forest community dominated by mixed conifers, principally balsam fir (Abies balsamea) and white spruce (Picea glauca) and scattered individuals of northern white cedar (Thuja occidentalis, or white pine (Pinus strobus), often with admixture of several deciduous species, most often trembling aspen (Populus tremuloides ), paper birch (Betula papyrifera), or red maple (Acer rubrum). Understory vegetation, both shrub and herb species, is well developed and diverse. Principle shrubs include beaked hazelnut (Corylus cornuta), speckled alder (Alnus incana) and red-osier dogwood (Cornus stolonifera). Herb layer typically is dominated by bracken fern (Pteridium aquilinum) and big-leaf aster (Aster macrophyllum). Other common species include wild strawberry (Fragaria virginiana), wild sarsaparilla (Aralia nudicaulis), bunchberry (Cornus Canadensis), snakeroot (Sanicula marilandica) and horsetails (Equisitum spp.). Depending on history of disturbance, two distinct community phases can be recognized.

Community 1.1
Mature Community

In the absence of stand replacing disturbance (period of 60-80 years) this community is dominated entirely by conifers, or it contains an admixture of old and decaying stems of aspen, and/or paper birch. This mixture is a result of typical succession process of conifer invasion of pioneer aspen-birch stands. In some areas, white pine, northern red cedar, or red maple are common associates. The tree reproduction layer is dominated by balsam fir, with lesser abundance of white spruce and/or red maple. The density of the reproduction is strongly dependent on the degree of canopy openings resulting from natural mortality, or small scale disturbance by wind and ice storms. The shrub and herb layers also depend on the degree of canopy opening. The dominant shrub typically is beaked hazel (Corylus cornuta). Common associates are speckled alder (Alnus incana), red osier dogwood (Cornus stolonifera) and wild rose (Rosa spp.). The herb layer often is well developed and species rich. Bracken fern (Pteridiun equilinum) and large-leaved aster (Aster macrophyllus) typically are most abundant herbs. Other common species include wild strawberry (Fragaria spp.), wild sarsaparilla (Aralia nudicaulis), bunchberry (Cornus Canadensis,) and horsetails (Equisitum spp.).

Community 1.2
Rejuvenated Community

Disturbance described in 1.1A typically removes some over-mature trees, especially old aspen and birch, from the overstory and releases advanced regeneration of balsam fir and white spruce and leads to community dominated by these species. Some presence of red maple and white pine may also result under favorable conditions, but trembling aspen and paper birch regeneration typically is not successful if disturbances do not include fire.

Pathway 1.1A
Community 1.1 to 1.2

A major canopy disturbance. Blow-down, severe ice storm, or large scale mortality in overstory due to old age.

Pathway 1.2A
Community 1.2 to 1.1

A long period without major canopy disturbances again leads to mature conifer community, similar to community phase 1.1, but without significant presence of aspen and birch.

State 2
Deciduous Forest State

Pure, or mixed, aspen – paper birch community replaces the reference state community 1.1. If seed source is present, red maple readily becomes member of this community.

Community 2.1
Aspen – Paper birch/Beaked hazelnut/Bracken fern-Big leaf aster

This deciduous forest state is composed of mixtures or pure stands of aspen and/or paper birch. Red maple may be present as well if the seed source is available.

State 3
Agriculture State

On clayey soils in MLRA 92 the most common agricultural activity is cultivation of hay.

Community 3.1
Cultivated crops primarily hay

This phase consists primarily of cleared agricultural land in hay production.

Transition T1A
State 1 to 2

Stand replacing disturbance that may include blow-down or ice storm, but must include fire to eliminate slash and competing vegetation and expose mineral soil to allow aspen and/or paper birch to colonize the site by seed. Alternatively, if the disrupted reference state community included aspen, the species may become re-established by vegetative means, which typically is more successful than colonization by seed.

Transition T1B
State 1 to 3

Removal of forest cover and tilling for agricultural crop production.

Restoration pathway R2
State 2 to 1

Deciduous forest community is slowly invaded by conifers (balsam fir, black spruce, white pine, white cedar).

Transition T2A
State 2 to 3

Removal of forest cover and tilling for agricultural crop production.

Restoration pathway R3A
State 3 to 1

Cessation of agricultural practices leads to natural reforestation, or site is replanted.

Restoration pathway R3B
State 3 to 2

Cessation of agricultural practices leads to natural reforestation, or site is replanted.

Additional community tables

Supporting information

Other references

Cleland, D.T.; Avers, P.E.; McNab, W.H.; Jensen, M.E.; Bailey, R.G., King, T.; Russell, W.E. 1997. National Hierarchical Framework of Ecological Units. Published in, Boyce, M. S.; Haney, A., ed. 1997. Ecosystem Management Applications for Sustainable Forest and Wildlife Resources. Yale University Press, New Haven, CT. pp. 181-200.

County Soil Surveys from Douglas, Bayfield, and Ashland Counties.

Curtis, J.T. 1959. Vegetation of Wisconsin: an ordination of plant communities. University of Wisconsin Press, Madison. 657 pp.

Davis, R.B. 2016. Bogs and Fens, A Guide to the Peatland Plants of Northeastern United States and Adjacent Canada. University Press of New England, Hanover and London. 296 pp.

Finley, R. 1976. Original vegetation of Wisconsin. Map compiled from U.S. General Land Office notes. U.S. Forest Service, North Central Forest Experiment Station, St. Paul, Minnesota.

Hvizdak, David. Personal knowledge and field experience.

Jahnke, J. and Gienccke, A. 2002. MLRA 92 Clay Till Field Investigations. Summary of field day investigations by Region 10 Soil Data Quality Specialists.

NatureServe. 2018. International Ecological Classification Satandard: Terrestrial Ecological Classifications. NautreServe Centreal Databases. Arlington, VA. U.S.A. Data current as of 28 August 2018.

Kotar, J. 1986. Soil – Habitat Type relationships in Michigan and Wisconsin. J. For. and Water Cons. 41(5): 348-350.

Kotar, J., J.A. Kovach and G. Brand. 1999. Analysis of the 1996 Wisconsin Forest Statistics by Habitat Type. U.S.D.A. For. Serv. N.C. Res. Stn. Gen. Tech. Rept. NC-207.

Kotar, J., J. A. Kovach, and T. L. Burger. 2002. A Guide to Forest Communities and Habitat Types of Northern Wisconsin. Second edition. University of Wisconsin-Madison, Department of Forest Ecology and Management, Madison.

Kotar, J., and T. L. Burger. 2017. Wetland Forest Habitat Type Classification System for Northern Wisconsin: A Guide for Land Managers and landowners. Wisconsin Department of Natural Resources, PUB-FR-627 2017, Madison.

Martin, L. 1965. The physical geography of Wisconsin. Third edition. The University of Wisconsin Press, Madison.

McNab, W.H. and P.W. Avers. 1994. Ecological Subregions of the United States: Section Descriptions. USDA For. Serv. Pun. WO-WSA-5, Washington, D.C.

Radeloff, V.C., D.J. Mladenoff, H.S. He and M.S. Boyce. 1999. Forest landscape change in Northwestern Wisconsin Pine Barrens from pre-European settlement to the present. Can. J. For. Res. 29: 1649-1659.

Schulte, L.A., and D.J. Mladenoff. 2001. The original U.S. public land sur¬vey records: their use and limitations in reconstructing pre-European settlement vegetation. Journal of Forestry 99:5–10.

Schulte, L.A., and D.J. Mladenoff. 2005. Severe wind and fire regimes in northern forests: historical variability at the regional scale. Ecology 86(2):431–445.

Soil Survey Staff. Input based on personal experience. Tim Miland, Scott Eversoll, Ryan Bevernitz, and Jason Nemecek.

Stearns, F. W. 1949. Ninety years change in a northern hardwood forest in Wisconsin. Ecology, 30: 350-58.

United States Department of Agriculture, Forest Service. 1989. Proceedings – Land Classification Based on Vegetation: Applications for Management. Gen. Tech. Report INT-527.

United States Department of Agriculture, Forest Service. 1990. Silvics of North America, Vol. 1, Hardwoods. Agricultural Handbook 654, Washington, D.C.

United States Department of Agriculture, Forest Service. 1990. Silvics of North America, Vol. 2, Conifers. Agricultural Handbook 654, Washington, D.C.

United States Department of Agriculture, Natural Resources Conservation Service. 2006. Land Resource and Major Land Resource Areas of the United Sates, the Caribbean, and the Pacific Basin. U.S. Department of Agriculture Handbook 296.

United States Department of Agriculture, Natural Resources Conservation Service. 2008. Hydrogeomorphic Wetland Classification System: An Overview and Modification to Better Meet the Needs of the Natural Resources Conservation Service. Technical Note No. 190-8-76. Washington D.C.

Wilde, S.A. 1933. The relation of soil and forest vegetation of the Lake States Region. Ecology 14: 94-105.

Wilde, S.A. 1976. Woodlands of Wisconsin. University of Wisconsin Cooperative Extension, Pub. G2780, 150 pp.

Wisconsin Department of Natural Resources. 2015. The ecological landscapes of Wisconsin: An assessment of ecological resources and a guide to planning sustainable management. Wisconsin Department of Natural Resources, PUB-SS-1131 2015, Madison.

Contributors

Jacob Prater, Associate Professor at University of Wisconsin Stevens Point
John Kotar, Ecological Specialist, independent contract
Bryant Scharenbroch, Assistant Professor at University of Wisconsin Stevens Point

Approval

Chris Tecklenburg, 4/09/2020

Acknowledgments

Contact for Lead Authors: Jacob Prater (jprater@uwsp.edu) Associate Professor at University of Wisconsin Stevens Point, John Kotar (jkotar@wsic.edu) Ecological Specialist, independent contract, and Bryant Scharenbroch Assistant Professor at University of Wisconsin Stevens Point

Rangeland health reference sheet

Interpreting Indicators of Rangeland Health is a qualitative assessment protocol used to determine ecosystem condition based on benchmark characteristics described in the Reference Sheet. A suite of 17 (or more) indicators are typically considered in an assessment. The ecological site(s) representative of an assessment location must be known prior to applying the protocol and must be verified based on soils and climate. Current plant community cannot be used to identify the ecological site.

Author(s)/participant(s)
Contact for lead author
Date	09/10/2022
Approved by	Chris Tecklenburg
Approval date
Composition (Indicators 10 and 12) based on	Annual Production