Ecological dynamics
This vegetation community on this PES is largely Northeastern Interior Dry Oak Forest, as classified by NatureServe. Hardwoods are most important in forested stands, but a Virginia/pitch pine component is present in places. Oak species characteristic of dry-mesic conditions (e.g., Quercus rubra, Quercus alba, Quercus velutina, and Quercus coccinea) and Carya spp. are dominant in mature stands. Quercus prinus may be present but is generally less important than the other oak species. Castanea dentata was a prominent tree before chestnut blight eradicated it as a canopy constituent.
NatureServe has also classified vegetation on this site in places as "Southern Ridge and Valley Dry Calcareous Forest". While there may be some species similarity, these sites cannot be classified as calcareous as soils range from slightly acidic throughout to extremely acid. Due to the variation in topography on this site and the range of slopes (0 - 80% in one case, 0 - 100% in another), microsite conditions can affect vegetation. In places, Appalachian cove forests might occur. The VDNH plots also indicate Montane Dry Calcareous Forest and Woodlands in one place. That plot fell on a Berks-Chiswell complex, 55 to 80 percent slopes mapunit. It is unclear whether or not that is due to a spatial error or perhaps is a correct classification of the community, which can occur on some calcareuous shale on interbedded geologies. It could also be due to an error in the soils map. Further investigation into the importance and abundance of this, more calcareous, plant community is needed on this site. For now, it will not be included in the PES description due to lack of data.
The Virginia Department of Natural Heritage plots on these soils fall under the following classifications: Rich Cove / Slope Forest, Montane Dry Calcareous Forest / Woodland, Eastern Hemlock - Hardwood Forest (not confident), and Oak / Heath Forest.
Rich Cove/Slope Forests are characterized by the VA-DNH as follows:
Characteristic trees include sugar maple (Acer saccharum), basswoods (Tilia americana var. americana and var. heterophylla), white ash (Fraxinus americana), tulip-tree (Liriodendron tulipifera), and yellow buckeye (Aesculus flava; chiefly south of the James River). Herbaceous growth is lush with spring ephemerals and leafy, shade-tolerant forbs such as blue cohosh (Caulophyllum thalictroides), yellow jewelweed (Impatiens pallida), large-flowered trillium (Trillium grandiflorum), wood-nettle (Laportea canadensis), common black cohosh (Actaea racemosa), sweet cicely (Osmorhiza claytonii), Virginia waterleaf (Hydrophyllum virginianum), large-leaf waterleaf (Hydrophyllum macrophyllum), large-flowered bellwort (Uvularia grandiflora), red trillium (Trillium erectum), yellow violets (Viola pubescens and Viola pensylvanica), white baneberry (Actaea pachypoda), two-leaved miterwort (Mitella diphylla), goat's-beard (Aruncus dioicus var. dioicus), yellow mandarin (Prosartes lanuginosa, = Disporum lanuginosum), showy skullcap (Scutellaria serrata), eastern blue-eyed-mary (Collinsia verna), Guyandotte beauty (Synandra hispidula), glade fern (Homalosorus pycnocarpos), and many others. Compositional variation related to substrate and elevation is complex but partitions convincingly into several major community types. The principal threats to rich cove forests are logging and invasion by shade-tolerant, non-native weeds, especially garlic-mustard (Alliaria petiolata).
References: Coulling and Rawinski (1999), Fleming (1999), Fleming and Coulling (2001), Fleming and Moorhead (1996), Fleming and Moorhead (2000), Johnson and Ware (1982), Olson and Hupp (1986), Rawinski et al. (1994), Rawinski et al. (1996), Rheinhardt and Ware (1984).
Full description: http://www.dcr.virginia.gov/natural_heritage/natural_communities/ncTIIa.shtml
Oak-Heath forests were described in two VA-DNH plots that fell on Weikert-Berks complex, 15 to 45 percent slopes
Weikert-Rock outcrop complex, 45 to 70 percent slopes mapunits. This group of oak-dominated forests is prominent on xeric, infertile upland sites in every physiographic province of Virginia, and is wide-ranging in the Appalachians and adjacent provinces outside of the Commonwealth. Fire plays an important role in this community. A full description can be found here: http://www.dcr.virginia.gov/natural_heritage/natural_communities/ncTIIIe.shtml
Because the VA-DNH was not confident about the classification on the one plot with Eastern Hemlock - Hardwood Forest, it will not be discussed in this PES description at this point in time. It may be revisited in the future, based on newly available information, including field visits.
A majority of this site has been converted to pasture/hay and cropland, especially on southern exposures. Principal crops are corn, wheat, oats, barley, Christmas trees and hay.
DeSelm had a few vegetation plots on Berks-Weikert soil map units in Union County, TN. He classified them as mixed mesophytic beech-buckeye-mixed oak, white oak-sugar maple, and beech-tulip poplar. This description does not match either NatureServe classification but does mesh well with the VDNH classification of Rich Cove/Slope Forest. That is probably the appropriate classification for this PES but could change with aspect. It is unclear whether or not the soil maps reflect change in aspect on this PES. That will have to be investigated in the field. There may be a dry forest component to this site that is important. DeSelm would have located his plots where there were the biggest trees so, his work might not correctly reflect the importance of aspect.
State 1
Mixed oak/hickory forest
In general, the reference community for these sites is mixed hardwood forests with oaks dominating. Species recorded on pedon description sheets for Berks and Weikert series include chestnut oak, hickory spp., northern red oak, white oak, and shortleaf pine.