Ecological dynamics
The vegetation groupings described in this section are based on the terrestrial ecological system classification and vegetation associations developed by NatureServe (Comer 2003) and the Natural Heritage Programs of Pennsylvania (Zimmerman et al. 2012), Virginia (Fleming et al. 2013), West Virginia (WVDNR 2014), and Maryland (Harrison 2004). Terrestrial ecological systems are specifically defined as a group of plant community types (associations) that tend to co-occur within landscapes with similar ecological processes, substrates, and/or environmental gradients. They are intended to provide a classification unit that is readily mappable, often from remote imagery, and readily identifiable by conservation and resource managers in the field. A given system will typically manifest itself in a landscape at intermediate geographic scales of tens to thousands of hectares and will persist for 50 or more years. A vegetation association is a plant community that is much more specific to a given soil, geology, landform, climate, hydrology, and disturbance history. It is the basic unit for vegetation classification. Each association will be named by the dominant species that occupy the different strata (tree, sapling, shrub, and herb). Within the NatureServe database, individual vegetation associations are assigned an identification number called a Community Element Global Code (CEGL). Most of the Information contained in this section was adapted from several sources, including the Nature Conservancy’s Northeast Terrestrial and Aquatic Habitat map (Anderson et al., 2013), NatureServe’s Ecological Systems of the United States (Comer 2003; NatureServe 2009), and Landfire’s Biophysical Settings and Existing Vegetation Type layers (Landfire 2010; Landfire 2013).
The characteristic forest system of this ecological site, and in most of the Northern Blue Ridge is the Northeastern Interior Dry-Mesic Oak Forest (NatureServe 2009; Anderson et. al 2013). This is an oak-dominated, mostly closed canopy forest that occurs as a matrix (dominant) type through much of the Appalachians. It occurs at low to mid elevations on gently rolling to steep topography on planar, slightly concave, and slightly convex slopes. Oak species characteristic of dry to mesic conditions and hickories are dominant in mature stands. These include Quercus rubra, Quercus alba, Quercus velutina, and Quercus coccinea (red, white, black, and scarlet oaks) and Carya spp. (hickories). Acer rubrum (Red maple), Betula lenta (Sweet birch), and Betula alleghaniensis (Yellow birch) may be common associates. Heath shrubs like Vaccinium spp. (blueberries) and Kalmia latifolia (Mountain laurel) are often present but not well developed. Castanea dentata (American chestnut) was a prominent tree before chestnut blight eradicated it as a canopy constituent.
Patches of exposed ridgetop and rocky areas with shallow soils may support a mosaic of open woodland, short-shrub or grassy herbaceous vegetation characteristic of the Southern and Central Appalachian Mafic Glade and Barrens system (NatureServe 2009; Anderson et. al 2013). Vegetation varies according to soil chemistry. Stunted and sparsely distributed tree species include Fraxinus Americana (white ash), Juniperus virginiana (Eastern red cedar), Quercus prinus (Chestnut oak), and Carya glabra (Pignut hickory). Rhus aromatic (aromatic sumac) and Physocarpus opulifolius (Ninebark) are common species in a shrub layer that may be thick. An herb layer dominated by graminoid species can be fairly dense away from bare rock; some typical forbs are Allium cernuum (nodding onion) and Helianthus divaricatus (woodland sunflower).
Concave slopes and drainageways will hold moisture and provide shade relative to other areas. Vegetation consists of forests dominated by various combinations of mesophytic (moisture-loving but non wetland) species of primarily deciduous trees. Liriodendron tulipifera (Tulip tree), Tilia Americana (American basswood), Fraxinus americana (White ash), Betula lenta (Sweet birch), PInus Strobus (Eastern white pine) and Tsuga canadensis (Eastern hemlock) are the most frequent dominant canopy species. These forests are characteristic of the Southern and Central Appalachian Cove Forest System (NatureServe 2009).
Disturbance agents in these forests include fire, wind throw, and ice damage. Gypsy moths can wreak havoc in the oak over story periodically. Oak forests historically have been maintained by periodic fire. Fire suppression since the early 20th century in the eastern United States is believed to be leading to the overall replacement of oaks with fire-sensitive, non-oak species like maples, beeches, birches, tulip poplars, and black cherry (Brose et. al., 2008). Oak forest regeneration is also hindered by heavy deer browsing (Latham et. al. 2005). Deer will selectively consume many native species including oak seedlings and acorns over less palatable species like hay-scented fern and several non-native species including Japanese barberry, Eurasian species of honeysuckle, and garlic mustard.
Much of this ecological site has been subjected to human activity including logging, settlement, or other disturbance, therefore many of the forests are mid successional, in which pines (typically Virginia or white) or tuliptree may be codominant or dominant. These ruderal (growing where the natural vegetation has been disturbed by humans) forests and woodlands comprise about 9% of the area and are generally characterized by unnatural combinations of species, primarily native species, though they often contain slight or substantial numbers and amounts of species alien to the region as well. Less than 5 percent of this ecological site has been converted to agricultural use, mainly pasture and hayland (Landfire 2013).
The information presented is representative of very complex vegetation communities. Key indicator plants and ecological processes are described to help inform land management decisions. Plant communities will differ across the major land resource region because of the naturally occurring variability in weather, soils, and aspect. The reference plant community is not necessarily the management goal. The species lists are representative and are not botanical descriptions of all species occurring, or potentially occurring, on this site. They are not intended to cover every situation or the full range of conditions, species, and responses for the site.
State 1
Reference
The reference state is a combination of several vegetation communities within the Northeastern Interior Dry-Mesic Oak Forest, the Southern and Central Appalachian Mafic Glade and Barrens, and the Southern and Central Appalachian Cove Forest Systems as defined by NatureServe (NatureServe 2009). These are oak-dominated, mostly closed canopy forests that occur as matrix (dominant) types through much of the Appalachians. Oak species characteristic of dry to mesic conditions and hickories are dominant in mature stands. Castanea dentata (American chestnut) was formerly an important canopy species prior to chestnut blight. Patches of exposed ridgetop and rocky areas with shallow soils may support a mosaic of open woodland, short-shrub or grassy herbaceous vegetation characteristic of glades and barrens. Coves and lower slopes support species that are more mesic (moisture loving) like maple, basswood, elm, birch, and beech. Vegetation varies according to soil chemistry. The reference communities listed below have been documented on this ecological site and are associated with the Northern Blue Ridge. Due to the heterogeneity and the broadness of this provisional ecological unit, they are not intended to cover every situation or the full range of conditions and species for this site. There are no transition pathways designated between the three communities in the reference state because the differences in vegetation are more controlled by landscape position than management or disturbance.
Community 1.1
Quercus rubra - Quercus prinus - Carya ovalis / (Cercis canadensis) / Solidago caesia Forest
The Northern Red Oak - Chestnut Oak - Red Hickory / (Eastern Redbud) / Wreath Goldenrod Forest, also known as the Central Appalachian Basic Oak Hickory Forest ((CEGL008514; NatureServe 2017), covers large areas at low to mid elevations in the Northern Blue Ridge and on some of its Piedmont foothills. It is generally associated with base-rich soils weathered from mafic igneous and metamorphic rocks. It also occurs less frequently on granitic rocks and calcareous metasiltstones and phyllites. This association is a true oak-hickory forest with mixed canopy dominance by several Quercus spp. (Oak species) and Carya spp. (Hickory species). In particular, Carya ovalis (Red hickory), Quercus rubra (Northern red oak), and Quercus prinus (Chestnut oak) are consistent codominants. Quercus alba (White oak), Quercus velutina (Black oak), Carya tomentosa (Mockernut hickory), Carya glabra (Pignut hickory), Fraxinus Americana (White ash), and Liriodendron tulipifera (Tuliptree) are less constant canopy species but achieve codominance in some stands. Carya spp., Quercus spp., Acer rubrum (Red maple), Nyssa sylvatica (Blackgum), Fraxinus Americana (White Ash), and Sassafras albidum (Sassafras) are well-represented in lower tree strata. Cercis canadensis (Eastern redbud) and, to a lesser extent, Cornus florida (Flowering dogwood) dominate the shrub and lowest tree layers, while Viburnum acerifolium (Mapleleaf viburnum) is a common low shrub. A large number of herbaceous species occur in the type.
Community 1.2
Acer saccharum - Tilia americana - Liriodendron tulipifera / Actaea racemosa Forest
The Sugar Maple - White Ash - American Basswood - Tuliptree / Black Baneberry Forest, also known as the Central Appalachian Rich Cove Forest (CEGL006237; NatureServe 2017), is a rich mesic, deciduous forest that occurs throughout the Central Appalachians as well as the Allegheny Plateau. Stands occur in coves and on lower slopes where soils are typically deep, fertile, and moderately to well-drained. The canopy is dominated by Acer saccharum (Sugar maple) with Fraxinus Americana (White ash), Liriodendron tulipifera (Tuliptree), and Tilia americana (American basswood) being very characteristic. Associated canopy trees include Quercus rubra (Northern red oak), Ostrya virginiana (hophornbeam), Ulmus rubra (Slippery elm), Acer rubrum (Red maple), Betula alleghaniensis (yellow birch), Betula lenta (Sweet birch), Fagus grandifolia (American Beech), Juglans nigra (Black walnut), Carya cordiformis (Bitternut hickory), and Prunus serotina (Black cherry). Some areas that are more acidic may have Pinus strobus (Eastern white pine), Tsuga Canadensis (Eastern hemlock), Quercus rubra (Northern red oak) and Quercus alba (White oak) as overstory dominants along with Tuliptree. The shrub layer is of variable composition, characterized by Cornus alternifolia (Alternateleaf dogwood), Hamamelis virginiana (American witchhazel), Asimina triloba (Pawpaw), Lonicera Canadensis (American fly honeysuckle), Rhododendron periclymenoides (Pink azalea), and Viburnum acerifolium (Mapleleaf viburnum). Lindera benzoin (Spice bush) is sparse to absent. The herb layer is diverse and made up of Adiantum pedatum (Northern maidenhair), Asarum canadense (Canadian wildginger), Actaea racemosa (Black baneberry), Cardamine spp. (Bittercress), Hepatica nobilis var. obtusa (Roundlobe hepatica), Hydrophyllum virginianum (Eastern waterleaf), Elymus hystrix (Eastern bottlebrush grass), Trillium grandiflorum (White trillium), Viola spp. (Violets), Dryopteris marginalis (Marginal woodfern), Botrychium virginianum, (Rattlesnake fern), Anemone quinquefolia (Wood anemone), Geranium maculatum (Spotted geranium), Caulophyllum thalictroides (Blue cohosh), Sanguinaria Canadensis (Bloodroot), Claytonia virginica (Virginia spring beauty), Allium tricoccum (Ramp), Cardamine concatenate (Cutleaf toothwort), Arisaema triphyllum (Jack in the pulpit), and Laportea Canadensis (Canadian woodnettle). The exotic weed Alliaria petiolata (Garlic mustard) is a rampant invader of some stands.
Community 1.3
Fraxinus americana / Physocarpus opulifolius/Carex pensylvanica - Allium cernuum - Wooded Grassland
The Sugar Maple - White Ash - American Basswood - Tuliptree / Black Baneberry Forest, also known as the Central Appalachian Rich Cove Forest (CEGL006237; NatureServe 2017), is a rich mesic, deciduous forest that occurs throughout the Central Appalachians as well as the Allegheny Plateau. Stands occur in coves and on lower slopes where soils are typically deep, fertile, and moderately to well-drained. The canopy is dominated by Acer saccharum (Sugar maple) with Fraxinus Americana (White ash), Liriodendron tulipifera (Tuliptree), and Tilia americana (American basswood) being very characteristic. Associated canopy trees include Quercus rubra (Northern red oak), Ostrya virginiana (hophornbeam), Ulmus rubra (Slippery elm), Acer rubrum (Red maple), Betula alleghaniensis (yellow birch), Betula lenta (Sweet birch), Fagus grandifolia (American Beech), Juglans nigra (Black walnut), Carya cordiformis (Bitternut hickory), and Prunus serotina (Black cherry). Some areas that are more acidic may have Pinus strobus (Eastern white pine), Tsuga Canadensis (Eastern hemlock), Quercus rubra (Northern red oak) and Quercus alba (White oak) as overstory dominants along with Tuliptree. The shrub layer is of variable composition, characterized by Cornus alternifolia (Alternateleaf dogwood), Hamamelis virginiana (American witchhazel), Asimina triloba (Pawpaw), Lonicera Canadensis (American fly honeysuckle), Rhododendron periclymenoides (Pink azalea), and Viburnum acerifolium (Mapleleaf viburnum). Lindera benzoin (Spice bush) is sparse to absent. The herb layer is diverse and made up of Adiantum pedatum (Northern maidenhair), Asarum canadense (Canadian wildginger), Actaea racemosa (Black baneberry), Cardamine spp. (Bittercress), Hepatica nobilis var. obtusa (Roundlobe hepatica), Hydrophyllum virginianum (Eastern waterleaf), Elymus hystrix (Eastern bottlebrush grass), Trillium grandiflorum (White trillium), Viola spp. (Violets), Dryopteris marginalis (Marginal woodfern), Botrychium virginianum, (Rattlesnake fern), Anemone quinquefolia (Wood anemone), Geranium maculatum (Spotted geranium), Caulophyllum thalictroides (Blue cohosh), Sanguinaria Canadensis (Bloodroot), Claytonia virginica (Virginia spring beauty), Allium tricoccum (Ramp), Cardamine concatenate (Cutleaf toothwort), Arisaema triphyllum (Jack in the pulpit), and Laportea Canadensis (Canadian woodnettle). The exotic weed Alliaria petiolata (Garlic mustard) is a rampant invader of some stands.
State 2
Post Logging - Fire Excluded Forest
Community 2.1
Liriodendron tulipifera - Quercus spp. Ruderal Forest
The Tuliptree – Oak Ruderal Forest (CEGL007221; NatureServe 2017) is a broadly defined successional community. These successional forests often follow cropping, clear-cut logging, or other severe disturbance, and are successional to mixed oak-hickory forests. Examples are common across large areas of the upland landscape which have previously been disturbed. Soils usually exhibit evidence of disturbance and may have little to no organic horizon development. Species found in stands attributable to this type may include a fairly diverse and varied composition. Acer rubrum (Red maple), Quercus spp. (Oak species), Betula lenta (Sweet birch), Oxydendrum arboretum (Sourwood), Acer saccharum (Sugar maple), and occasionally Liquidambar styraciflua (Sweetgum), Ilex opaca (American holly), or Robinia pseudoacacia (Black locust) may be common in stands of this type. Where oaks are present, they are frequently multi-stemmed, resulting from coppicing. The conifer Tsuga Canadensis (Eastern hemlock) is abundant in the understories of some stands. Shrub composition is variable but may include Sambucus nigra ssp. Canadensis (American black elderberry), Rhododendron maximum (Great laurel), Hamamelis virginiana (American witchhazel), and Vaccinium pallidum (Blue Ridge blueberry). Herbs are likewise variable and may include Dioscorea quaternata (Four leaf yam), Lysimachia quadrifolia (Whorled yellow loosestrife), Maianthemum racemosum (Feathery false lily of the valley), Solidago curtisii (Mountain decumbent goldenrod), Symphyotrichum prenanthoides (Crookedstem aster), Polystichum acrostichoides (Christmas fern), Dryopteris intermedia (Intermediate woodfern), Arisaema triphyllum ssp. Triphyllum (Jack in the pulpit), Packera aurea (Golden ragwort), Amphicarpaea bracteata (American hogpeanut), Thelypteris noveboracensis (New York fern), Lycopodium digitatum (fan clubmoss), and Geranium maculatum (Spotted geranium). Oak forests historically have been maintained by periodic fire. Fire suppression since the early 20th century in the eastern United States is believed to be leading to the overall replacement of oaks with fire-sensitive, non-oak species like maples, beeches, birches, tulip poplars, and black cherry (Brose et. al., 2008). Historic logging has resulted in areas of even-aged tree stands.
State 3
Agricultural - Pasture
Community 3.1
Dactylis glomerata - Festuca spp. - Solidago canadensis Ruderal Mesic Meadow
The Orchardgrass - Fescue - Canada Goldenrod Ruderal Mesic Meadow Alliance (A1190, NatureServe 2017) is a broadly defined community which includes mesic abandoned pastures and agricultural fields and is largely composed of non-native cool-season grasses and herbs (generally of European origin) in the early stages of succession. Species composition varies from site to site, depending on land-use history and perhaps soil type, but in general this vegetation is quite wide-ranging in northeastern and midwestern states. Dominant grasses vary from site to site but generally include the exotic grasses Agrostis stolonifera (Creeping bentgrass), Agrostis hyemalis (Winter bentgrass), Anthoxanthum odoratum, (Sweet vernalgrass), Bromus inermis (Smooth Brome), Bromus tectorum (Cheatgrass), Dactylis glomerata (Orchardgrass), Schedonorus arundinaceum (Tall fescue), Lolium perenne (Perennial ryegrass), Phleum pretense (Timothy) as well as weedy natives such as Elymus repens (Quackgrass), Poa pratensis (Kentucky bluegrass), and, less commonly, Schizachyrium scoparium (Little bluestem). Herbaceous species may be minor or dominant and include various Solidago spp. (goldenrods), Sympyotrichum spp. (Asters), and other native and non-native species. Less than 5% of the Mixed Metamorphic and Granitic Upland provisional ecological site is in pasture and hayland (Landfire 2013).
State 4
Transitional Invaded Woodland or Forest
Community 4.1
Prunus serotina - Liriodendron tulipifera - Acer rubrum - (Robinia pseudoacacia) Ruderal Forest
The Black Cherry - Tuliptree - Red Maple - White Ash - (Black Locust) Ruderal Forest (CEGL006599; NatureServe 2017) is an early-successional woody vegetation community of the northeastern United States that occurs on sites that are becoming reforested after having been cleared for agriculture or otherwise heavily modified in the past. This modified successional forest has been documented on the Mixed Metamorphic and Granitic Uplands ecological site and analysis of existing vegetation data from Landfire shows that approximately 9% of this ecological site is a ruderal hardwood and conifer forest. Characteristics of this community are highly variable ranging from closed forest, to woodland, to open to dense shrubland. Tree species often include some combination of Prunus serotina (Black cherry), Liriodendron tulipifera (Tuliptree), Fraxinus Americana (White ash), Robinia pseudoacacia (Black locust), and Acer rubrum (Red maple). Other associates can include Juglans nigra (Black walnut), Sassafras albidum (Sassafras), Betula populifolia (Gray birch), Juniperus virginiana (Eastern redcedar), Acer negundo (Boxelder), Acer saccharinum (Silver maple), Ailanthus altissima (Tree of heaven), Ulmus Americana (American elm), Quercus spp. (Oak species), Betula lenta (Sweet birch), Amelanchier spp. (Serviceberry), Pinus strobus (Eastern white pine), and Populus grandidentata (Bigtooth aspen). The low-shrub layer, if present, is usually characterized by the presence of Rubus spp. (Blackberry). This layer is often dominated by exotic species such as Lonicera tatarica (Tatarian honeysuckle), Lonicera morrowii (Morrow’s honeysuckle), Rhamnus cathartica (Common buckthorn), Crataegus spp.(Hawthorn), Rosa multiflora (Multiflora rose), and Berberis thunbergii (Japanese barberry). The herbaceous layer is variable, often containing grasses and forbs of both native and exotic origin. Common species include Ageratina altissima var. altissima (White snakeroot), Polygonum persicaria (Spotted ladysthumb), Impatiens capensis (Jewelweed), Glechoma hederacea (Ground ivy), Polystichum acrostichoides (Christmas fern), Calystegia sepium ssp. Sepium (Hedge false bindweed), Galium aparine (Stickywilly), Oxalis stricta (Common yellow oxalis), Polygonum virginianum (Jumpseed), Dennstaedtia punctilobula (Eastern hayscented fern), Arisaema triphyllum (Jack in the pulpit), Allium vineale (Wild garlic), and Veronica officinalis (Common gypsyweed), among many others. The invasive species Alliaria petiolate (Garlic mustard), Microstegium vimineum (Nepalese browntop), and Polygonum caespitosum (Oriental lady’s thumb) can be abundant in this disturbed forest type. These forests are often young and resulted from the colonization of old agricultural fields by woody species. Recent disturbance or abundant invasive species give these forest stands a weedy character. It is unlikely that these stands will succeed to a natural plant community dominated by native species.
Transition T1A
State 1 to 2
Logging, but no agricultural conversion. Trees are allowed to stump sprout, soil is minimally disturbed, seed bank remains. Fire suppression allows fire sensitive species like tuliptree, red maple, and birches to out compete oak seedlings in the understory.
Transition T1C
State 1 to 3
Logging, clearing, and then planting of non-native pasture grass mixes, and grazing. Maintenance with periodic mowing to prevent trees and shrubs from reestablishing.
Transition T1B
State 1 to 4
Logging followed by agricultural conversion. Soil surface is disturbed by tillage or clearing of tree stumps and vegetation, and allows colonization by successional species. Field is then abandoned. If surrounding forests are still intact, they can provide native seed sources. If surrounding forests are not intact, or area is surrounded by agriculture, or other human development, nonnative species may become dominant.
Restoration pathway R2A
State 2 to 1
Control of understory to allow oak seedling recruitment. Prescribed fire will further advance the growth of oaks over fire sensitive species.
Transition T2A
State 2 to 3
Logging, clearing, and then planting of non-native pasture grass mixes, and grazing. Maintenance with periodic mowing to prevent trees and shrubs from reestablishing.
Transition T2B
State 2 to 4
Logging followed by agricultural conversion and then abandonment. Soil surface is disturbed by tillage or clearing of tree stumps and vegetation, and allows colonization by successional species. If surrounding forests are still intact, they can provide native seed sources. If surrounding forests are not intact, or area is surrounded by agriculture, or other human development, nonnative species may become dominant.
Restoration pathway R3A
State 3 to 2
Return to the reference or post logged minimally managed state may require a very long term series of costly management options and stages. Many species may need to be planted or seeded to restore the system. If using acorns, direct seeding must be done fairly heavily. Herbivory can be a problem as well as competition from faster growing species. Depending on the existing seed bank and the proximity of a mature forest from which to recruit seeds, ruderal forests may regain a mixed forest stand. Nevertheless, sites that have been cleared and tilled have significant soil disturbance which may include compaction, erosion, loss of native soil structure, loss of soil organic matter, disruption of soil microorganisms, all which affect the soil’s nutrient availability and water holding capacity (Duiker and Myers, 2005). These characteristics favor recolonization by plant species that have wind dispersed seeds (verses those that propagate through underground roots called rhizomes, or which have heavy seeds that stay near the parent tree), are shade intolerant, have rapid to moderate growth rates, and drought tolerance. These communities are distinctly different from the reference forest state (Dyer, 2010).
Transition T3A
State 3 to 4
Abandonment of pasture or old field. Discontinue mowing and do not allow grazing. Allow natural regeneration.
Restoration pathway R4A
State 4 to 1
Return to the reference or post logged minimally managed state may require a very long term series of costly management options and stages. Many species may need to be planted or seeded to restore the system. If using acorns, direct seeding must be done fairly heavily. Herbivory can be a problem as well as competition from faster growing species. Depending on the existing seed bank and the proximity of a mature forest from which to recruit seeds, ruderal forests may regain a mixed forest stand. Nevertheless, sites that have been cleared and tilled have significant soil disturbance which may include compaction, erosion, loss of native soil structure, loss of soil organic matter, disruption of soil microorganisms, all which affect the soil’s nutrient availability and water holding capacity (Duiker and Myers, 2005). These characteristics favor recolonization by plant species that have wind dispersed seeds (verses those that propagate through underground roots called rhizomes, or which have heavy seeds that stay near the parent tree), are shade intolerant, have rapid to moderate growth rates, and drought tolerance. These communities are distinctly different from the reference forest state (Dyer, 2010).
Restoration pathway R4B
State 4 to 2
Return to the reference or post logged minimally managed state may require a very long term series of costly management options and stages. Many species may need to be planted or seeded to restore the system. If using acorns, direct seeding must be done fairly heavily. Herbivory can be a problem as well as competition from faster growing species. Depending on the existing seed bank and the proximity of a mature forest from which to recruit seeds, ruderal forests may regain a mixed forest stand. Nevertheless, sites that have been cleared and tilled have significant soil disturbance which may include compaction, erosion, loss of native soil structure, loss of soil organic matter, disruption of soil microorganisms, all which affect the soil’s nutrient availability and water holding capacity (Duiker and Myers, 2005). These characteristics favor recolonization by plant species that have wind dispersed seeds (verses those that propagate through underground roots called rhizomes, or which have heavy seeds that stay near the parent tree), are shade intolerant, have rapid to moderate growth rates, and drought tolerance. These communities are distinctly different from the reference forest state (Dyer, 2010).
Transition T4A
State 4 to 3
Logging, clearing, and then planting of non-native pasture grass mixes, and grazing. Maintenance with periodic mowing to prevent trees and shrubs from establishing.