Ecological dynamics
**Disclaimer** This PROVISIONAL ecological site was developed for an extended MLRA concept where the foothills (10-14" precipitation Frigid climatic zone) was included in this site. This has created an exaggerated or elevated production value for this site description.
Potential vegetation on this site is dominated by mid cool-season perennial grasses. Other significant vegetation includes winterfat, black and big sagebrush, rubber rabbitbrush, juniper, and a variety of forbs. The expected potential composition for this site is about 75% grasses, 10% forbs and 15% woody plants. The composition and production will vary naturally due to historical use, fluctuating precipitation and fire frequency.
As this site deteriorates, species such as threadleaf sedge, blue grama, big and/or black sagebrush, and juniper will increase. Plains pricklypear and weedy annuals will invade. Mid cool season grasses such as Griffiths and bluebunch wheatgrass, Indian ricegrass, needleandthread, and rhizomatous wheatgrasses will decrease in frequency and production.
A mixture of shrubs may become dominant on some areas with an absence of fire. Wildfires are actively controlled in recent times so chemical control using herbicides has replaced the historic role of fire on this site. Recently, prescribed burning has regained some popularity.
Due to the amount and pattern of the precipitation, the shrub component typically is not resilient once it has been removed if a healthy and vigorous stand of grass exists and is maintained. The exception to this is where the herbaceous component is severely degraded at the time of treatment, growing conditions are unfavorable after treatment, and/or recovery periods are inadequate.
The Historic Climax Plant Community (description follows the plant community diagram) has been determined by study of rangeland relic areas, or areas protected from excessive disturbance. Trends in plant communities going from heavily grazed areas to lightly grazed areas, seasonal use pastures, and historical accounts have also been used.
The following is a State and Transition Model Diagram that illustrates the common plant communities (states) that can occur on the site and the transitions between these communities. The ecological processes will be discussed in more detail in the plant community narratives following the diagram.
Plant Community Narratives
Following are the narratives for each of the described plant communities. These plant communities may not represent every possibility, but they probably are the most prevalent and repeatable plant communities. The plant composition tables shown above have been developed from the best available knowledge at the time of this revision. As more data is collected, some of these plant communities may be revised or removed, and new ones may be added. None of these plant communities should necessarily be thought of as “Desired Plant Communities”. According to the USDA NRCS National Range and Pasture Handbook, Desired Plant Communities (DPC’s) will be determined by the decision-makers and will meet minimum quality criteria established by the NRCS. The main purpose for including any description of a plant community here is to capture the current knowledge and experience at the time of this revision.
State 1
Bluebunch wheatgrass/ Indian Ricegrass
Community 1.1
Bluebunch wheatgrass/ Indian Ricegrass
The interpretive plant community for this site is the Historic Climax Plant Community. This state evolved with grazing by large herbivores and periodic fires. Potential vegetation is about 75% grasses or grass-like plants, 10% forbs, and 15% woody plants. Cool season mid-grasses dominate this state. The major grasses include Griffiths and bluebunch wheatgrass, Indian ricegrass needleandthread, and rhizomatous wheatgrass. Other grasses occurring in the state include prairie junegrass, Sandberg bluegrass, bottlebrush squirreltail, red threeawn, blue grama, and threadleaf sedge. Winterfat, and black sagebrush are conspicuous components of this site but other shrubs occur including big sagebrush, juniper, and rubber rabbitbrush. Antelope bitterbrush will occur on sites associated with the upper limits of this precipitation zone. A variety of forbs also occurs in this state and plant diversity is high (see Plant Composition Table).
The total annual production (air-dry weight) of this state is about 400 pounds per acre, but it can range from about 250 lbs./acre in unfavorable years to about 600 lbs./acre in above average years.
The state is stable and well adapted to the Northern Intermountain Desertic Basins climatic conditions. The diversity in plant species allows for high drought resistance. This is a sustainable plant community (site/soil stability, watershed function, and biologic integrity).
Transitions or pathways leading to other plant communities are as follows:
• Moderate, Continuous Season-Long grazing will convert the plant community to the Perennial Grass/Mixed Shrub Plant Community.
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State 2
Perennial Grass/Mixed Shrub
Community 2.1
Perennial Grass/Mixed Shrub
Historically, this plant community evolved under grazing by large ungulates and a low fire frequency. Currently, it is found under moderate, season-long grazing by livestock, and will be exacerbated by prolonged drought conditions. In addition, the fire regime for this site has been modified and extended periods without fire is now common. Shrubs and forbs make up an increasingly significant portion of this plant community. Cool-season grasses make up the majority of the understory with the balance made up of short warm-season grasses.
The dominant grasses include Griffiths and bluebunch wheatgrass, needleandthread, rhizomatous wheatgrasses, Sandberg bluegrass, red threeawn, threadleaf sedge, and blue grama. Forbs commonly found in this plant community include scarlet globemallow, Hood’s phlox, sulfur flower buckwheat, and lemon scurfpea. Black and big sagebrushes, rubber rabbitbrush and juniper can make up to 25% of the total annual production. Plains pricklypear will also occur.
When compared to the Historic Climax Plant Community, Griffiths and bluebunch wheatgrass and Indian ricegrass have decreased. Shrubs and warm season grasses/grass-likes such as threadleaf sedge, blue grama, and red threeawn have increased.
The total annual production (air-dry weight) of this state is about 320 pounds per acre, but it can range from about 200 lbs./acre in unfavorable years to about 450 lbs./acre in above average years.
This plant community is resistant to change. The herbaceous species present are well adapted to grazing; however, species composition can be altered through long-term overgrazing. The herbaceous component is mostly intact and plant vigor and replacement capabilities are sufficient. Water flow patterns and litter movement may be occurring but only on steeper slopes. Incidence of pedestalling is minimal. Soils are mostly stable and the surface shows minimum soil loss. The watershed is functioning and the biotic community is intact.
Transitions or pathways leading to other plant communities are as follows:
• Prescribed grazing or possibly long-term prescribed grazing will convert this plant community to the HCPC. The probability of this occurring is high especially if rotational grazing along with short deferred grazing is implemented as part of prescribed method of use. In addition, the removal of fire suppression will allow a somewhat natural fire regime to reoccur to more easily transition between this plant community and the HCPC. A prescribed fire treatment can be useful to hasten this transition if desired.
• Frequent and severe grazing plus no fires will convert the plant community to the Mixed Shrub/Bare Ground Plant community. The probability of this occurring is high. This is especially evident on areas where drought or heavy browsing does not adversely impact the shrub stand.
• Frequent and severe grazing (yearlong) plus brush management, will convert the plant community to a Threadleaf Sedge and/or Blue Grama Sod/ Bare Ground Plant community. The probability for is high especially on areas were the shrubs have been heavily browsed or removed by natural or human causes. Drought can also exasperate this transition.
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State 3
Mixed Shrub/ Bare Ground
Community 3.1
Mixed Shrub/ Bare Ground
This plant community is the result of frequent and severe grazing and protection from fire. Black and big sagebrushes, rubber rabbitbrush, and juniper dominate this plant community as the annual production of shrubs exceeds 25%. Shrubs, forbs, and warm season grasses comprise the significant components of the plant community and the preferred cool season grasses have been greatly reduced.
The dominant grasses are prairie junegrass, Sandberg bluegrass, red threeawn, threadleaf sedge, and blue grama. Fringed sagewort, stemless goldenweed, basin rayless daisy, and sulfur flower buckwheat are some of the dominant forbs. Weedy annual species such cheatgrass, kochia, Russian thistle, halogeton and a variety of mustards may occupy the site. Cactus has increased. Plant diversity is moderate to poor. The interspaces between plants have expanded leaving the amount of bare ground more prevalent. When compared with the HCPC, the annual production is less as the perennial mid cool season grasses have been reduced, but the shrub production compensates for some of the decline in the herbaceous production.
The total annual production (air-dry weight) of this state is about 275 pounds per acre, but it can range from about 100 lbs./acre in unfavorable years to about 350 lbs./acre in above average years.
This plant community is resistant to change as the stand becomes more decadent. These areas may actually be more resistant to fire as less fine fuels are available and the bare ground between the shrubs is increased. The herbaceous component is not as diverse and plant vigor and species regeneration capabilities of cool-season perennials are deficient. The removal of grazing does not seem to affect the plant composition or structure of the plant community.
Soil erosion is accelerated because of increased bare ground. Water flow patterns and pedestalling are obvious. Infiltration is reduced and runoff is increased. Rill channels may be noticeable in the interspaces and gullies may be establishing where rills have concentrated down slope.
Transitions or pathways leading to other plant communities are as follows:
• Brush management, followed by prescribed grazing and seeding, will return this plant community to at or near the HCPC. If prescribed fire is used as a means to reduce or remove the shrubs, sufficient fine fuels will need to be present. This may require deferment from grazing prior to treatment. Post management is critical to ensure success. This can range from two or more years of rest to partial growing season deferment, depending on the condition of the understory at the time of treatment and the growing conditions following treatment. In the case of an intense wildfire that occurs when desirable plants are not completely dormant, the length of time required to reach the HCPC may be increased and seeding of natives is recommended.
• Brush management, followed by frequent and severe grazing, will convert the plant community to a Threadleaf Sedge and/or Blue Grama Sod/ Bare Ground Plant community. The probability of this occurring is high, because of the amount of bare ground exposed to weedy annuals and sod formers.
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State 4
Threadleaf Sedge and/ or Blue Grama Sod
Community 4.1
Threadleaf Sedge and/ or Blue Grama Sod
This plant community is created when the Perennial Grass/Mixed Shrub Plant Community is subjected to severe heavy yearlong grazing and the shrub component has been removed by heavy browsing, wildfire or human means. Additionally, this plant community can occur as a result of the Mixed Shrub/Bare Ground Plant Community being subjected to fire or brush management and not followed by prescribed grazing. Weedy annuals, threadleaf sedge, and/or blue grama are the most dominant plants. Weedy annuals occupy any open bare ground areas, while threadleaf sedge and/or blue grama can form extensive sod patches. Rubber rabbitbrush may or may not be present on the site as this species may quickly re-establish the site after a fire. However, heavy browsing by large ungulates will significantly reduce or remove this shrub. Shadscale saltbush can also be found on this site and become more prominent.
Compared to the HCPC, weedy annual species are widespread and may include cheatgrass, kochia, Russian thistle, halogeton and a variety of mustards. Cactus and sageworts have invaded significantly. Noxious weeds such as Russian knapweed, leafy spurge, or Canada thistle may invade the site if a seed source is available. Virtually all other cool-season mid-grasses are absent or severely decreased. Blue grama and threadleaf sedge have significantly increased from what is found in the HCPC. Shrubs have been removed with the exception of patches of rubber rabbitbrush and black sagebrush. Plant diversity is low.
The total annual production (air-dry weight) of this state is about 80 pounds per acre, but it can range from about 35 lbs./acre in unfavorable years to about 120 lbs./acre in above average years.
This plant community is relatively stable and resistant to overgrazing. Annuals and sod forming grasses are effectively competing against the establishment of perennial cool-season grasses. Plant diversity is greatly altered and the herbaceous component is not intact. Recruitment of perennial grasses is not occurring and the replacement potential is absent. The biotic integrity is missing.
On areas with a well established sod plant community, water infiltration will be significantly affected. While this sod protects the area itself, adjacent on-site and off-site areas are impacted by excessive runoff that can cause rill channels and gully erosion. Water flow patterns and pedestalling are obvious. The watershed may or may not be functional.
Transitions or pathways leading to other plant communities are as follows:
• Grazing land mechanical treatment (chiseling, etc.) and pricklypear cactus control (if needed), followed by prescribed grazing, will return this plant community to near Historic Climax Plant Community condition. The sod areas are extremely resistant to change and will require grazing land mechanical treatments, such as chiseling to revert to a more preferred state. This may not be possible given the presence of gravels and cobbles on the soil surface, which can also exclude reseeding the area. If applicable, mechanical treatments and reseeding native plant species are recommended. This should be followed by proper grazing management to accelerate recovery where few desirable plants remain.
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