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Ecological site R058BY178WY

Wetland (WL) 10-17” PZ

Last updated: 9/15/2024
Accessed: 11/21/2024

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General information

Provisional. A provisional ecological site description has undergone quality control and quality assurance review. It contains a working state and transition model and enough information to identify the ecological site.

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Figure 1. Mapped extent

Areas shown in blue indicate the maximum mapped extent of this ecological site. Other ecological sites likely occur within the highlighted areas. It is also possible for this ecological site to occur outside of highlighted areas if detailed soil survey has not been completed or recently updated.

MLRA notes

Major Land Resource Area (MLRA): 058B–Northern Rolling High Plains, Southern Part

MLRA 58B is located in northeastern Wyoming (95 percent) and extreme southeastern Montana (5 percent). It is comprised of sedimentary plains, scoria hills, and river valleys. The major rivers include the Powder, Tongue, Belle Fourche, Cheyenne, and North Platte. Tributaries include the Little Powder River, Little Missouri River, Clear Creek, Crazy Woman Creek, and others. This MLRA is traversed by Interstates 25 and 90, and U.S. Highways 14 and 16. The extent of MLRA 58B covers approximately 12.3 million acres. Major land uses include rangeland (approximately 93 percent), cropland, pasture, and hayland (approximately 2 percent), and forest, urban, and miscellaneous uses (approximately 5 percent). Cities include Buffalo, Casper, Sheridan, and Gillette, WY. Land ownership is mostly private. Federal lands include the Thunder Basin National Grassland (U.S. Forest Service) and lands administered by the Bureau of Land Management. Areas of interest in MLRA 58B in Wyoming include Fort Phil Kearny State Historic Site, Glendo State Park, and Lake DeSmet. The elevations in MLRA 58B increase gradually from north to south and range from approximately 2,900 to 5,900 feet. A few buttes are higher than 6,800 feet. The average annual precipitation in this area ranges from 10 to 17 inches per year. Precipitation occurs mostly during the growing season, often during rapidly developing thunderstorms. Mean annual air temperature is 46 degrees Fahrenheit. Summer temperatures may exceed 100 degrees Fahrenheit. Winter temperatures may drop to below zero. Snowfall averages 45 inches per year, but varies from 25 to over 70 inches in some locales.

Classification relationships

USDA Natural Resources Conservation Service (NRCS):
Land Resource Region─G Western Great Plains Range and Irrigation; Major Land Resource Area (MLRA)─ 58B Northern Rolling High Plains, Southern Part (USDA, 2006).

Relationship to Other Classifications:

USDA Forest Service (FS) Classification Hierarchy:

Province─331 Great Plains-Palouse Dry Steppe; Section─331G-Powder River Basin; Subsections─331Gb Montana Shale Plains, 331Ge Powder River Basin, 331Gf South Powder River Basin-Scoria Hills (Cleland et al, 1997)

Environmental Protection Agency (EPA) Classification Hierarchy:

Level III Ecoregion─43 Northwestern Great Plains; Level IV Ecoregion─43p Scoria Hills, 43q Mesic-Dissected Plains, 43w Powder River Basin (EPA, 2013)
https://www.epa.gov/eco-research/ecoregions

Ecological site concept

The Wetland 10-17” Precipitation Zone (PZ) site occurs where a seasonal or perennial water table exists of sedimentary plains or lowlands; on stream terraces, drainageways, and oxbows; adjacent to streams, springs, seeps and sloughs. It is a mixed-grass prairie, with warm- and cool season rhizomatous grasses, and cool-season grass-likes, and (sedges, rushes, and bulrushes), and cool-season bunchgrasses, followed by a minor component of forbs and shrubs.

Associated sites

R058BY142WY	Saline Subirrigated (SS) 10-14” PZ Saline Subirrigated occurs generally in close partnership with wetland sites, as the initial ring or next step in the transition of the water table depth and influence on salt burdened sites.
R058BY130WY	Overflow (Ov) 10-14” PZ Overflow is the highest on the landform/landscape of this complex. Overflow is generally drier with less water influence than any other site.
R058BY174WY	Subirrigated (Sb) 10-17” PZ Subirrigated occurs generally in close partnership with wetland sites, as the initial ring or next step in the transition of the water table depth and influence on the site.
R058BY128WY	Lowland (LL) 10-14” PZ Lowland occurs more closely to subirrigate or overflow, and is generally slightly above wetlands and subirrigated sites in the landform/landscape.

Similar sites

R058BY178WY	Wetland (WL) 10-17” PZ Subirrigated (Sb) is less productive.
R058BY142WY	Saline Subirrigated (SS) 10-14” PZ Saline Subirrigated (SS) has more salt-tolerant species and is less productive.

R058BY178WY

Wetland (WL) 10-17” PZ

Subirrigated (Sb) is less productive.

R058BY142WY

Saline Subirrigated (SS) 10-14” PZ

Saline Subirrigated (SS) has more salt-tolerant species and is less productive.

Table 1. Dominant plant species

Tree	Not specified
Shrub	Not specified
Herbaceous	Not specified

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Physiographic features

This site occurs on nearly level stream terraces, drainageways, and oxbows; on sedimentary plains or lowlands, adjacent to springs, seeps, and sloughs.

Table 2. Representative physiographic features

Landforms	(1) Stream terrace (2) Drainageway (3) Oxbow (4) Flood plain (5) Playa
Runoff class	Negligible to high
Flooding duration	Very brief (4 to 48 hours) to brief (2 to 7 days)
Flooding frequency	Occasional to frequent
Ponding frequency	None
Elevation	3,500 – 5,400 ft
Slope	6%
Water table depth	18 in
Aspect	Aspect is not a significant factor

Climatic features

The average annual precipitation ranges from 10 to 17 inches per year across MLRA 58B. There are two precipitation zones (PZ). The 10 to 14 inch precipitation zone is predominant across the MLRA, including portions of Sheridan, Johnson, and Natrona Counties; portions of Campbell and Converse Counties; and smaller portions of Weston and Niobrara Counties. The 15 to 17 inch precipitation zone occurs in northern and eastern portions of the MLRA, including portions of Sheridan, Campbell, and western Crook Counties. Wide fluctuations in precipitation may occur from year to year, and occasional periods of extended drought (longer than one year in duration) can be expected. Two-thirds of the annual precipitation occurs during the growing season from May through September. Mean Annual Air Temperature (MAAT) is 46 degrees Fahrenheit. Cold air outbreaks from Canada in winter move rapidly from northwest to southeast and account for extreme minimum temperatures. Chinook winds may also occur in winter and bring rapid rises in temperature. Extreme storms may occur during the winter, but most severely affect ranching operations during late winter and spring. High-intensity afternoon thunderstorms may occur during the summer. Annual wind speeds average about 5 mph. Daytime winds are generally stronger than nighttime winds. Occasional strong storms may bring brief periods of high winds with gusts of more than 75 mph. The average length of the freeze-free period (28 degrees Fahrenheit) is 125 days and generally occurs from May 16 to September 19. The average frost-free period (32 degrees Fahrenheit) is 101 days and generally occurs from June 1 to September 9.

The growth of native cool-season plants begins in late April to early May with peak growth occurring in mid to late June. Native warm-season plants begin growth in late May to early June and continue into August. Regrowth of cool-season plants occurs in September in most years, depending upon moisture.

Note: The climate described here is based on historic climate station data and is averaged to provide an overview of the annual precipitation, temperatures, and growing season. Future climate is beyond the scope of this document. However, research to determine the effects of elevated CO2 and heating on mixed-grass prairie ecosystems, and how it may relate to future plant communities, is ongoing.

For detailed information, or to find a specific climate station, visit the Western Regional Climate Center (WRCC) website: Western Regional Climate Center, Historical Data, Western U.S. Climate summaries, NOAA Coop Stations, Wyoming (Note: Montana climate stations are also listed under the Wyoming link).
https://wrcc.dri.edu/summary/Climsmwy.html

Wind speed averages can be found at the WRCC home page, under the Specialty Climate tab: https://wrcc.dri.edu/

The following tables represent area-wide climate data for the 10 to 14 inch precipitation zone:

Table 3. Representative climatic features

Frost-free period (characteristic range)	88-105 days
Freeze-free period (characteristic range)	122-129 days
Precipitation total (characteristic range)	13-15 in
Frost-free period (actual range)	84-110 days
Freeze-free period (actual range)	118-130 days
Precipitation total (actual range)	10-17 in
Frost-free period (average)	101 days
Freeze-free period (average)	125 days
Precipitation total (average)	14 in

Characteristic range

Actual range

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Figure 2. Monthly precipitation range

Characteristic range

Actual range

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Figure 3. Monthly minimum temperature range

Characteristic range

Actual range

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Figure 4. Monthly maximum temperature range

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Figure 5. Monthly average minimum and maximum temperature

Figure 6. Annual precipitation pattern

Figure 7. Annual average temperature pattern

Climate stations used

(1) DULL CTR 1SE [USC00482725], Douglas, WY
(2) MIDWEST [USC00486195], Midwest, WY
(3) SHERIDAN CO AP [USW00024029], Sheridan, WY
(4) LEITER 9N [USC00485506], Clearmont, WY
(5) DOUGLAS 1 SE [USC00482685], Douglas, WY
(6) GILLETTE 4SE [USC00483855], Gillette, WY
(7) DILLINGER [USC00482580], Gillette, WY
(8) GLENROCK 5 ESE [USC00483950], Glenrock, WY
(9) BUFFALO [USC00481165], Buffalo, WY
(10) WRIGHT 12W [USC00489805], Gillette, WY

Influencing water features

Wetland 10-14” PZ ecological site: Map units in this ESD typically have hydric components for stream terraces, drainageways, and oxbows that have a water table.

Note: The water table in some areas is artificially induced, caused by seepage from nearby irrigation ditches, canals, or reservoirs.

Wetland description

System - Palustrine
Subsystem - None
Class - Emergent Wetland
Subclass - Persistent

Stream Type: C (Rosgen)

Note: This is a general overview for the site concept and is not a wetland determination.

Soil features

The soils on Wetland 10 to 17 inch Precipitation Zone ecological site are deep to very deep, poorly drained and formed in alluvium derived from shale and interbedded siltstone. The permeability class ranges from slow to moderate for the soils on this site. The available water capacity is typically high but may range to very high in some soils. The surface layer of the soils in this site are typically clay loam or loam but may include silt loam, silty clay loam, silty clay, or clay. The surface layer ranges from a depth of 4 to 10 inches thick. The subsoil is typically clay loam, silty clay loam, or clay. There are typically 0 to 5 percent rock fragments in the subsoil. Soils in this site are typically calcareous to the surface, but some pedons are leached as deep as 13 to 24 inches. These soils are susceptible to erosion by water and wind. The hazard for water erosion accelerates with increasing slope. The soil moisture regime is aquic. The soil temperature regime is mesic.

Major soil series correlated to this ecological site include: None listed.

The attributes listed represent 0-40 inches in depth or to the first restrictive layer.

Table 4. Representative soil features

Parent material	(1) Alluvium – shale (2) Alluvium – siltstone
Surface texture	(1) Clay loam (2) Loam (3) Silt loam (4) Silty clay loam (5) Silty clay (6) Clay
Drainage class	Very poorly drained to poorly drained
Permeability class	Slow to moderate
Soil depth	20 – 80 in
Available water capacity (Depth not specified)	2.8 – 8.8 in
Calcium carbonate equivalent (Depth not specified)	5%
Electrical conductivity (Depth not specified)	4 mmhos/cm
Sodium adsorption ratio (Depth not specified)	5
Soil reaction (1:1 water) (Depth not specified)	7.4 – 8.4
Subsurface fragment volume <=3" (Depth not specified)	5%

Ecological dynamics

The Reference state is the plant community in which interpretations are primarily based and is used as a reference in order to understand the original potential of the site. The Reference state evolved under the combined influences of climatic conditions, periodic fire activity, grazing by large herbivores, and impacts from small mammals and insects. Changes may occur to the Reference state due to management actions such as continuous season-long or year-long grazing, increased stocking rates, climatic conditions such as drought, and natural events. The Reference state is characterized by warm-season rhizomatous tallgrass, cool-season rhizomatous grass, and grass-like plants. A minor component of grass-likes such as mountain rush (also known as Baltic rush) and other rush, spikerush, and bulrush species; forbs and shrubs are also present. Noxious weeds such as purple loosestrife may invade. The Reference state is not necessarily the management goal, as other vegetative states may be considered desired plant communities as long as critical resource concerns are met.

Recurrent drought has historically impacted the vegetation of this region. Changes in species composition and production will vary, depending upon the duration and severity of the drought cycle, and prior grazing management. Long-term water table alteration or drainage may cause a complete disruption of the hydrologic function and biotic integrity resulting in the crossing of an ecological threshold and the replacement of the natural states and plant communities applicable to the site. Likewise, artificially-induced water tables, (i.e. reservoirs, irrigation canals, etc.), are not considered the central concept of the wetland ecological site but occur on the landscape.

As the Wetland 10-17” PZ ecological site begins to shift from a combination of frequent and severe defoliation during the growing season, species such as spike sedge and mountain rush (Baltic rush), will increase forming a cool-season-dominated plant community. Cattails will increase. Mountain rush will eventually become sod-bound. Noxious weeds will also invade. Once these events have occurred, it is difficult for native perennial plants to reestablish. There are various transitional stages which may occur on this ecological site. The information presented is representative of a dynamic set of plant communities that illustrate the complex interaction of several ecological processes.

State and transition model

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Ecosystem states

1. Reference

2. Sod Bound

3. Increased Bare Ground

T1A	-	Excessive grazing without adequate recovery periods, or frequent and severe defoliation
T2A	-	Long-term excessive grazing or frequent and severe defoliation without adequate recovery between grazing events, or heavy, excessive grazing with overstocking

State 1 submodel, plant communities

1.1. Narrowleaf Willow, Prairie Cordgrass, and Nebraska Sedge

1.2. Narrowleaf Willow, Prairie Cordgrass, and Bluejoint Reedgrass

1.1A	-	Non-use or lack of fire
1.2A	-	The return of grazing with adequate recovery or normal fire frequency

State 2 submodel, plant communities

2.1. Reed Canarygrass, Mountain Rush, and Broadleaf Cattail

State 3 submodel, plant communities

3.1. Mountain Rush, Barnyardgrass, Smartweed

State 1
Reference

The Reference state is characterized by two distinct plant communities. The plant communities, and various successional stages between them, represent the natural range of variability within the Reference state.

Dominant plant species

narrowleaf willow (Salix exigua), shrub
prairie cordgrass (Spartina pectinata), grass
Nebraska sedge (Carex nebrascensis), grass
bluejoint (Calamagrostis canadensis), grass

Community 1.1
Narrowleaf Willow, Prairie Cordgrass, and Nebraska Sedge

This is the interpretive plant community for the Wetland 10 to 17 inch Precipitation Zone ecological site. This community developed with grazing by large herbivores and is suited to grazing by domestic livestock. Historically, fires likely occurred infrequently, and were randomly distributed. This plant community can be found on areas where grazed plants receive adequate periods of recovery during the growing season. The potential vegetation is about 80 percent grasses and grass-likes, 5 to 10 percent forbs, and 5 to 10 percent woody plants. The major grasses and grass-likes include prairie cordgrass, bluejoint and northern reedgrasses, and Nebraska sedge. Minor species include slender and western wheatgrasses, basin wildrye, and switchgrass; other minor species are tufted hairgrass, foxtail barley, reed canarygrass, and native bluegrasses. A minor component of grass-likes such as mountain rush (also known as Baltic rush), inland and spike sedges, and spikerush and bulrush species. Forbs such as horsetail, arrowgrass, iris, blue-eyed grass, and American licorice. Other species include water hemlock and a minor component of shrubs such as willows are present. In the Wetland 10 to 17 inch Precipitation Zone (PZ) ecological site, the total annual production (air-dry weight) is about 5,300 pounds per acre during an average year, but it can range from about 4,500 pounds per acre in unfavorable years to about 6,000 pounds per acre in above-average years. Defoliation levels should be determined as part of a grazing management plan based on objectives. Community dynamics (nutrient and water cycles, and energy flow) are functioning properly. Infiltration rates are moderate, and soil erosion is low. Litter is properly distributed where vegetative cover is continuous. Decadence and natural plant mortality are low. This community is resistant to many disturbances except excessive grazing, tillage, or development into urban or other uses.

Figure 8. Annual production by plant type (representative values) or group (midpoint values)

Table 5. Annual production by plant type

Plant type	Low (lb/acre)	Representative value (lb/acre)	High (lb/acre)
Grass/Grasslike	6100	7550	9000
Forb	400	500	600
Shrub/Vine	500	550	600
Total	7000	8600	10200

Figure 9. Plant community growth curve (percent production by month). WY1403, 10-14NP free water sites.

Jan	Feb	Mar	Apr	May	Jun	Jul	Aug	Sep	Oct	Nov	Dec
J	F	M	A	M	J	J	A	S	O	N	D
0	0	0	5	15	30	25	10	15	0	0	0

Community 1.2
Narrowleaf Willow, Prairie Cordgrass, and Bluejoint Reedgrass

This plant community developed under the absence of grazing and/or fire. Plant species resemble community 1.1; however, frequency and production will be reduced. Standing dead canopy may prevent sunlight from reaching plant crowns. Much of the available nutrients are tied up in standing dead plant material and litter. Eventually, litter levels can become high enough to cause decadence or mortality of the stand. Bunchgrasses, such as slender wheatgrass, typically develop dead centers and rhizomatous grasses can form small decadent communities due to a lack of impact by grazing animals. The absence of animal traffic to break down litter will slow nutrient recycling. Water flow patterns and pedestalling can become apparent. Infiltration is reduced and runoff is increased. In advanced states of non-use (rest) or lack of fire, bare areas will increase, causing an erosion concern. Total annual production can vary substantially.

Figure 10. Plant community growth curve (percent production by month). WY1403, 10-14NP free water sites.

Jan	Feb	Mar	Apr	May	Jun	Jul	Aug	Sep	Oct	Nov	Dec
J	F	M	A	M	J	J	A	S	O	N	D
0	0	0	5	15	30	25	10	15	0	0	0

Pathway 1.1A
Community 1.1 to 1.2

Non-use or lack of fire will cause community 1.1 to shift toward community 1.2. Plant decadence and standing dead plant material will impede energy flow. Initially, excess litter will increase. Eventually, native plant density begins to decrease and weeds and introduced species may begin to invade. Water and nutrient cycles will be impaired as a result of this community pathway.

Pathway 1.2A
Community 1.2 to 1.1

The return of grazing with adequate recovery or normal fire frequency can shift this plant community toward community 1.1. This change can occur in a relatively short time frame with the return of these disturbances.

State 2
Sod Bound

This state is characterized by the sod-bound nature of the plant community. An ecological threshold has been crossed and a significant amount of production and diversity has been lost when compared to the Reference state. Significant biotic and soil changes have negatively impacted energy flow, and nutrient and hydrologic cycles. This is a very stable state, resistant to change due to the high tolerance of inland saltgrass to grazing, the development of a shallow root system (root pan), and subsequent changes in hydrology and nutrient cycling. The loss of other functional/structural groups such as cool-season bunch and rhizomatous grasses, forbs, and shrubs, reduces the biodiversity productivity of this site. This alternative state should be tested through long-term observation of ecosystem behavior and repeated application of conservation and restoration practices. This state should be reevaluated and refined.

Dominant plant species

reed canarygrass (Phalaris arundinacea), grass
arctic rush (Juncus arcticus), grass
broadleaf cattail (Typha latifolia), other herbaceous

Community 2.1
Reed Canarygrass, Mountain Rush, and Broadleaf Cattail

This plant community develops under long-term frequent and severe defoliation, without adequate recovery between grazing events. Prairie cordgrass, northern reedgrass, and Nebraska sedge have been removed. Low-growing unpalatable sedges and cattails have increased. Mountain rush (also known as Baltic rush) persists in a sod-bound condition. Willows are present near the drier edges of the plant community. The plant community lacks diversity and is resistant to change. Energy flow and the water and mineral cycles have been impaired due to the loss of tallgrass species and deep-rooted forbs and shrubs. Soil compaction can be a concern if continuously grazed during wet cycles. Litter levels are very low and unevenly distributed. The total annual production (air-dry weight) is about 3,000 pounds per acre during an average year, but it can range from about 2,000 pounds per acre in unfavorable years to about 4,000 pounds per acre in above-average years.

Figure 11. Plant community growth curve (percent production by month). WY1403, 10-14NP free water sites.

Jan	Feb	Mar	Apr	May	Jun	Jul	Aug	Sep	Oct	Nov	Dec
J	F	M	A	M	J	J	A	S	O	N	D
0	0	0	5	15	30	25	10	15	0	0	0

State 3
Increased Bare Ground

The Increased Bare Ground state develops with heavy, excessive grazing and/or excessive defoliation. An ecological threshold has been crossed. The Increased Bare Ground state denotes changes in infiltration, runoff, aggregate stability and species composition. The changes in water movement and the plant community affect changes in hydrologic functionality, biotic integrity, and soil and site stability. Infiltration, runoff, and soil erosion vary depending on the vegetation present. Erosion and loss of organic matter and carbon reserves are resource concerns. Desertification is advanced. This alternative state should be tested and refined in future updates through long-term observation of ecosystem behavior, and repeated application of conservation and restoration practices.

Dominant plant species

arctic rush (Juncus arcticus), grass
cockspur grass (Echinochloa), grass
knotweed (Polygonum), other herbaceous

Community 3.1
Mountain Rush, Barnyardgrass, Smartweed

This plant community occurs where the rangeland is grazed year-round, at high stock densities. Physical impact such as trampling, soil compaction, and trailing typically contribute to this transition. Mountain rush still dominates the plant community. However increased bare ground results in a broken, sod-bound appearance. Bare ground may continue to increase if water table levels are low. Baltic rush, smartweed, American licorice, and cattails dominate this plant community. Noxious weeds such as purple loosestrife may invade. Forage palatability for livestock is low. This plant community is highly variable in both species composition and production. Average annual production should be determined on-site. This plant community is very resistant to change because of the lack of native species and the amount of invasive species present. Wind and water erosion may occur if bare ground has increased. Litter amounts are greatly reduced. Continued heavy use will cause severe compaction problems.

Figure 12. Plant community growth curve (percent production by month). WY1403, 10-14NP free water sites.

Jan	Feb	Mar	Apr	May	Jun	Jul	Aug	Sep	Oct	Nov	Dec
J	F	M	A	M	J	J	A	S	O	N	D
0	0	0	5	15	30	25	10	15	0	0	0

Transition T1A
State 1 to 2

Excessive grazing without adequate recovery periods, or frequent and severe defoliation, will shift this plant community across an ecological threshold toward the Sod Bound state. Biotic integrity and hydrologic function will be impaired because of this transition.

Transition T2A
State 2 to 3

Long-term excessive grazing or frequent and severe defoliation without adequate recovery between grazing events, or heavy, excessive grazing with overstocking will cause a shift across an ecological threshold to the Increased Bare Ground state. Erosion and loss of organic matter, along with invasion of introduced plants and noxious weeds, are resource concerns.

Additional community tables

Table 6. Community 1.1 plant community composition

Group	Common name	Symbol	Scientific name	Annual production (lb/acre)	Foliar cover (%)
Grass/Grasslike
1	Cool-Season Tallgrasses			740–1740
	bluejoint	CACA4	Calamagrostis canadensis	1000–1500	5–25
	northern reedgrass	CASTI3	Calamagrostis stricta ssp. inexpansa	1000–1500	5–25
	tufted hairgrass	DECE	Deschampsia cespitosa	1000–1500	5–25
	basin wildrye	LECI4	Leymus cinereus	400–600	5–10
2	Warm-Season Tallgrass			500–1500
	prairie cordgrass	SPPE	Spartina pectinata	400–600	1–10
3	Grass-likes			500–1000
	Nebraska sedge	CANE2	Carex nebrascensis	1000–1500	5–25
	inland sedge	CAIN11	Carex interior	400–600	1–10
	spike sedge	CANA2	Carex nardina	400–600	1–10
	sedge	CAREX	Carex	400–600	1–10
	rush	JUNCU	Juncus	400–600	1–10
	bulrush	SCHOE6	Schoenoplectus	400–600	1–10
	spikerush	ELEOC	Eleocharis	400–600	1–10
	mountain rush	JUARL	Juncus arcticus ssp. littoralis	400–600	1–10
Forb
5	Forbs			230–480
	broadleaf cattail	TYLA	Typha latifolia	200–300	1–5
	Forb, perennial	2FP	Forb, perennial	200–300	1–5
	aster	ASTER	Aster	200–300	1–5
	water hemlock	CICUT	Cicuta	200–300	1–5
	horsetail	EQUIS	Equisetum	200–300	1–5
	American licorice	GLLE3	Glycyrrhiza lepidota	200–300	1–5
	wild mint	MEAR4	Mentha arvensis	200–300	1–5
	Rocky Mountain iris	IRMI	Iris missouriensis	200–300	1–5
	Pennsylvania smartweed	POPE2	Polygonum pensylvanicum	200–300	1–5
	poison hemlock	COMA2	Conium maculatum	200–300	1–5
	blue-eyed grass	SISYR	Sisyrinchium	200–300	1–5
	dock	RUMEX	Rumex	200–300	1–5
	seaside arrowgrass	TRMA20	Triglochin maritima	200–300	1–5
	narrowleaf cattail	TYAN	Typha angustifolia	200–300	1–5
Shrub/Vine
6	Shrubs			250–500
	willow	SALIX	Salix	500–600	1–5
	Shrub (>.5m)	2SHRUB	Shrub (>.5m)	500–600	1–5
	Subshrub (<.5m)	2SUBS	Subshrub (<.5m)	500–600	1–5

Interpretations

Animal community

Animal Community – Wildlife Interpretations (from 2001 ESD; will be revised in future updates)

Nebraska Sedge/ Northern Reedgrass (Reference): The predominance of grasses in this plant community favors grazers and mixed-feeders, such as bison, elk, and antelope. Suitable thermal and escape cover for deer may be limited due to the low quantities of woody plants. This plant community may provide brood-rearing and foraging areas for sage grouse. Other birds that would frequent this plant community include red-wing blackbirds, sandhill cranes, Wilson snipe, western meadowlarks, and golden eagles. Many small mammals would occur here.

Baltic Rush/Cattail Plant Community: This plant community may be useful for the same large grazers that would use the Reference Plant Community. However, the plant community composition is less diverse, and thus less apt to meet the seasonal needs of these animals. It may provide some foraging opportunities for sage grouse when it occurs proximal to woody cover. Good grasshopper habitat equals good foraging for birds.

Baltic rush/Smartweed Plant Community: This plant community may be useful for the same large grazers that would use the Reference Plant Community. However, the plant community composition is less diverse, and thus less apt to meet the seasonal needs of these animals. It may provide some foraging opportunities for sage grouse when it occurs proximal to woody cover. Grasshopper habitat provides good foraging for birds.

Animal Community – Grazing Interpretations (updated in the 2019 Provisional revision)

The following table is a guide to stocking rates for the plant communities described in the Wetland 10 to 17 inch Precipitation Zone ecological site. These are conservative estimates for initial planning. On-site conditions will vary, and stocking rates should be adjusted based on range inventories, animal kind and class, forage availability (adjusted for slope, distance to water), and the type of grazing system (number of pastures, planned moves, etc.), all of which is determined in the conservation planning process.

The following stocking rates are based on the total annual forage production in a normal year multiplied by 25 percent harvest efficiency of preferred and desirable forage species, divided by 912 pounds of ingested air-dry vegetation for an animal unit per month (Natl. Range and Pasture Handbook, 1997). An animal unit month is defined as the amount of forage required by one livestock animal, with or without one calf, for one month, and is shortened to AUM.

Example:
5,000 lbs. per acre X 25% Harvest Efficiency = 1,250 lbs. forage demand for one month. 1,250 lbs. per acre/912 demand per AUM = 1.4

Plant Community (PC) Production (total lbs./acre in a normal year) and Stocking Rate (AUM/acre) are listed below:

Reference Plant Community 4000-6000 3.0
Baltic rush/Cattail 2000-4000 2.0
Baltic rush/Smartweed 1200-2500 1.0

Increased Bare Ground PC (*) (*)

* Highly variable stocking rates must be determined on-site.
Grazing by domestic livestock is one of the major income-producing industries in the area. Rangelands in this area provide year-long forage under prescribed grazing for cattle, sheep, horses, and other herbivores. During the dormant period, livestock may need supplementation based on reliable forage analysis.

Hydrological functions

Climate is the principal factor limiting forage production on this site. This site is dominated by soils in hydrologic groups B and C, with localized areas in hydrologic group D. Infiltration and runoff potential for this site varies from moderate to high depending upon soil hydrologic group and water table. Runoff will be high on this site because the soil may be saturated. (Refer to Part 630, NRCS National Engineering Handbook for detailed hydraulic information.

Rills and gullies should not typically be present. Water flow patterns should be barely distinguishable if at all present. Litter should fall in place. Fine litter will move after average to high rainfall events. Litter does not travel far typically being trapped in small bunches by the extensive vegetative cover. Litter movement may be extensive after major runoff or flooding events. Chemical and physical crusts are rare to non-existent. Cryptogamic crusts are present, but only cover 1 to 2 percent of the soil surface.

Recreational uses

This site provides hunting opportunities for upland game species. The wide variety of plants which bloom from spring until fall have an esthetic value that appeals to visitors.

Wood products

No appreciable wood products are present on the site.

Other products

None noted.

Other information

Site Development & Testing Plan

General Data (MLRA and Revision Notes, Hierarchical Classification, Ecological Site Concept, Physiographic, Climate, and Water Features, and Soils Data):

Updated. All “Required” items complete to Provisional level.

Community Phase Data (Ecological Dynamics, STM, Transition & Recovery Pathways, Reference Plant Community, Species Composition List, Annual Production Table):

Updated. All “Required” items complete to Provisional level.

Annual Production Table is from the “Previously Approved” ESD (2001).

The Annual Production Table and Species Composition List will be reviewed for future updates at the Approved level.

Each Alternative State/Community:

Complete to Provisional level.

Supporting Information (Site Interpretations, Assoc. & Similar Sites, Inventory Data References, Agency/State Correlation, References):

Updated. All “Required” items complete to Provisional level.

Wildlife Interpretations: Narrative is from “Previously Approved” ESD (2001). Wildlife species will need to be updated at the next Approved level.

Livestock Interpretations: Plant community names and stocking rates updated.

Hydrology, Recreational Uses, Wood Products, and Other Products carried over from previously “Approved” ESD (2001).

Existing NRI Inventory Data References updated. More field data collection is needed to support this site concept.

Reference Sheet:

Rangeland Health Reference Sheet carried over from previously “Approved” ESD (2005).
It will be updated at the next “Approved” level.

“Future work, as described in a project plan, to validate the information in this provisional ecological site description is needed. This will include field activities to collect low and medium intensity sampling, soil correlations, and analysis of that data. Annual field reviews should be done by soil scientists and vegetation specialists. A final field review, peer review, quality control, and quality assurance reviews of the ESD will be needed to produce the final document.” (NI 430_306 ESI and ESD, April 2015)

Supporting information

Inventory data references

Inventory data has been collected on private and federal lands by the following methods:
• Double Sampling (Determining Vegetation Production and Stocking Rates, WY-ECS-1)
• Rangeland Health (Interpreting Indicators of Rangeland Health, Version 4, 2005)
• Soil Stability (Interpreting Indicators of Rangeland Health, Version 4, 2005)
• Line Point Intercept (Monitoring Manual for Grassland, Shrubland, and Savanna Ecosystems, Volume II, 2005)
• Soil Pedon Descriptions (Field Book for Describing and Sampling Soils, Version 3, 2012)
• SCS-RANGE-417 (Production & Composition Record for Native Grazing Lands)

National Resources Inventory (NRI)
Number of Records: 0
Sample Period: 2005-2017
Counties:

USDA - Agricultural Research Service (ARS)
Thunder Basin National Grassland
Plant Community Responses to Historical Wildfire in a Shrubland/Grassland Ecotone
• Number of Records: 140
• Sample Period: 2014-2021
• State: Wyoming
• Counties: Campbell, Converse, Crook, Niobrara, and Weston

Additional data collection includes ESI data collection in conjunction with Soil Surveys conducted within MLRA 58B; ocular estimates; rangeland vegetative clipping for NRCS program support; field observations from experienced rangeland personnel

Data collection for this ecological site was done in conjunction with the progressive soil surveys within MLRA 58B Northern Rolling High Plains (Southern Part)

Note: Revisions to soil surveys are on-going. For the most recent updates, visit the Web Soil Survey, the official site for soils information: http://websoilsurvey.nrcs.usda.gov/app/WebSoilSurvey.aspx

Other references

Anderson, R.C. 2006. Evolution and origin of the central grassland of North America: Climate, fire, and mammalian grazers. Journal of the Torrey Botanical Society 133:626–647.

Bragg, T.B. 1995. The physical environment of the Great Plains grasslands. In: A. Joern and K.H. Keeler (eds) The changing prairie. Oxford University Press, Oxford, UK, pages 49–81.

Branson, D.H. and G.A. Sword. 2010. An experimental analysis of grasshopper community responses to fire and livestock grazing in a northern mixed-grass prairie. Environmental Entomology 39:1441–1446.

Brinson, M.M. 1993. A hydrogeomorphic classification for wetlands. Technical Report WRP–DE–4. U.S. Army Corps of Engineers Waterways Experiment Station, Vicksburg, MS.

Cleland, D., P. Avers, W.H. McNab, M. Jensen, R. Bailey, T. King, and W. Russell. 1997. National hierarchical framework of ecological units. In: Ecosystem Management: Applications for Sustainable Forest and Wildlife Resources, Yale University Press.

Coupland, R.T. 1958. The effects of fluctuations in weather upon the grasslands of the Great Plains. Botanical Review 24:273–317.

Davis, S.K., R.J. Fisher, S.L. Skinner, T.L. Shaffer, and R.M. Brigham. 2013. Songbird abundance in native and planted grassland varies with type and amount of grassland in the surrounding landscape. Journal of Wildlife Management 77:908–919.

DeLuca, T.H. and P. Lesica. 1996. Long-term harmful effects of crested wheatgrass on Great Plains grassland ecosystems. Journal of Soil and Water Conservation 51:408–409.

Derner, J.D. and R.H. Hart. 2007. Grazing-induced modifications to peak standing crop in northern mixed-grass prairie. Rangeland Ecology and Management 60:270–276.

Derner, J.D. and A.J. Whitman. 2009. Plant interspaces resulting from contrasting grazing management in northern mixed-grass prairie: Implications for ecosystem function. Rangeland Ecology and Management 62:83–88.

Derner, J.D., W.K. Lauenroth, P. Stapp, and D.J. Augustine. 2009. Livestock as ecosystem engineers for grassland bird habitat in the western Great Plains of North America. Rangeland Ecology and Management 62:111–118.

Dillehay, T.D. 1974. Late Quaternary bison population changes on the southern Plains. Plains Anthropologist 19:180–196.

Dormaar, J.F. and S. Smoliak. 1985. Recovery of vegetative cover and soil organic matter during revegetation of abandoned farmland in a semiarid climate. Journal of Range Management 38:487–491.

Guyette, Richard P., M.C. Stambaugh, D.C. Dey, and R.M. Muzika. (2012). Predicting fire frequency with chemistry and climate. Ecosystems, 15: 322-335.

Harmoney, K.R. 2007. Grazing and burning Japanese brome (Bromus japonicus) on mixed grass rangelands. Rangeland Ecology and Management 60:479–486.

Heitschmidt, R.K., and L.T. Vermeire. 2005. An ecological and economic risk avoidance drought management decision support system. In: J.A. Milne (ed.) Pastoral systems in marginal environments, 20th International Grasslands Congress, July, 2005. Page 178.

Knopf, F.L. 1996. Prairie legacies—Birds. In: F.B. Samson and F.L. Knopf (eds.) Prairie conservation: Preserving North America’s most endangered ecosystem. Island Press, Washington, DC. Pages 135–148.

Knopf, F.L., and F.B. Samson. 1997. Conservation of grassland vertebrates. In: F.B. Samson and F.L. Knopf (eds.) Ecology and conservation of Great Plains vertebrates: Ecological Studies 125. Springer-Verlag, New York, NY. Pages 273–289.

Lauenroth, W.K., O.E. Sala, D.P. Coffin, and T.B. Kirchner. 1994. The importance of soil water in recruitment of Bouteloua gracilis in the shortgrass steppe. Ecological Applications 4:741–749.

Laycock, W.A. 1988. History of grassland plowing and grass planting on the Great Plains. In: J.E. Mitchell (ed.) Impacts of the Conservation Reserve Program in the Great Plains—symposium proceedings, September 16–18, 1987. U.S. Dept. of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station, General Technical Report RM-158.

Malloch, D.W., K.A. Pirozynski, and P.H. Raven. 1980. Ecological and evolutionary significance of mycorrhizal symbioses in vascular plants (a review). Proceedings of the National Academy of Sciences 77:2113–2118.

Ogle, S.M., W.A. Reiners, and K.G. Gerow. 2003. Impacts of exotic annual brome grasses (Bromus spp.) on ecosystem properties of the northern mixed grass prairie. American Midland Naturalist 149:46–58.

Roath, L.R. 1988. Implications of land conversions and management for the future. In: J.E. Mitchell (ed.) Impacts of the Conservation Reserve Program in the Great Plains—symposium proceedings, September 16–18, 1987. U.S. Dept. of Agriculture, Forest Service, Rocky Mountain Forest and Range Experiment Station, General Technical Report RM-158.

Smoliak, S. and J.F. Dormaar. 1985. Productivity of Russian wildrye and crested wheatgrass and their effect on prairie soils. Journal of Range Management 38:403–405.

Smoliak, S., J.F. Dormaar, and A. Johnston. 1972. Long-term grazing effects on Stipa-Bouteloua prairie soils. Journal of Range Management 25:246–250.

Soil Survey Division Staff. 2017. Soil survey manual. U.S. Dept. of Agriculture Handbook 18.

Soil Survey Staff. Official Soil Series Descriptions. U.S. Dept. of Agriculture, Natural Resources Conservation Service. Available online. https://www.nrcs.usda.gov/wps/portal/nrcs/detail/soils/home/?cid=nrcs142p2_053587 Accessed 15 November, 2017.

Soil Survey Staff. Soil Survey Geographic (SSURGO) database. U.S. Dept. of Agriculture, Natural Resources Conservation Service.

Soil Survey Staff. 2014. Keys to Soil Taxonomy, 12th edition. U.S. Dept. of Agriculture, Natural Resources Conservation Service.
Soil Survey Staff. 2018. Web Soil Survey. U.S. Dept. of Agriculture, Natural Resources Conservation Service. Available online. https://websoilsurvey.nrcs.usda.gov/app/. Accessed 15 February, 2018.

Soller, D.R. 2001. Map showing the thickness and character of Quaternary sediments in the glaciated United States east of the Rocky Mountains. U.S. Geological Survey Miscellaneous Investigations Series I-1970-E, scale 1:3,500,000.

Stewart, Omer C. 2002. Forgotten Fires. Univ. of Oklahoma Press, Norman, OK.

United States Department of Agriculture, Natural Resources Conservation Service. Glossary of landform and geologic terms. National Soil Survey Handbook, Title 430-VI, Part 629.02c. Available online. http://www.nrcs.usda.gov/wps/portal/nrcs/detail/soils/ref/?cid=nrcs142p2_054242. Accessed 16 January, 2018.

United States Army Corps of Engineers. 1987. Corps of Engineers wetlands delineation manual. Wetlands Research Program Technical Report Y-87-1. Available online. http://www.lrh.usace.army.mil/Portals/38/docs/USACE%2087%20Wetland%20Delineation%20Manual.pdf. Waterways Experiment Station, Vicksburg, MS.

United States Environmental Protection Agency, National Health and Environmental Effects Research Laboratory. 2013. Level III ecoregions of the continental United States. Available online. https://www.epa.gov/eco-research/ecoregions Accessed 30 January, 2019.

United States Department of Agriculture, Natural Resources Conservation Service. 2010a. Field indicators of hydric soils in the United States, version 7.0.

United States Department of Agriculture, Natural Resources Conservation Service. 2013a. Climate data. National Water and Climate Center. Available online. http://www.wcc.nrcs.usda.gov/climate. Accessed 13 October, 2017.

United States Department of Agriculture, Natural Resources Conservation Service. 2006. Land resource regions and major land resource areas of the United States, the Caribbean, and the Pacific Basin. Agriculture Handbook 296.

United States Department of Agriculture, Natural Resources Conservation Service. 2013b. National Soil Information System. Available online. https://www.nrcs.usda.gov/wps/portal/nrcs/detail/soils/survey/geo/?Cid=nrcs142p2_053552. Accessed 30 October, 2017.

United States Department of the Interior, Geological Survey. 2008. LANDFIRE 1.1.0 Vegetation Dynamics Models. http://landfire.cr.usgs.gov/viewer/.

United States Department of the Interior, Geological Survey. 2011. LANDFIRE 1.1.0 Existing Vegetation Types. http://landfire.cr.usgs.gov/viewer/.

Willeke, G.E. 1994. The national drought atlas [CD ROM]. U.S. Army Corps of Engineers, Water Resources Support Center, Institute for Water Resources Report 94-NDS-4.

Wilson, S.D. and J.M. Shay. 1990. Competition, fire, and nutrients in a mixed-grass prairie. Ecology 71:1959–1967.

With, K.A. 2010. McCown’s longspur (Rhynchophanes mccownii). In: A. Poole (ed.) The birds of North America [online]. Cornell Lab of Ornithology, Ithaca, NY. https://birdsna.org/Species-Account/bna/home.

Augustine, D.J., J. Derner, D. Milchunas, D. Blumenthal, and L. Porensky. 2017. Grazing moderates increases in C3 grass abundance over seven decades across a soil texture gradient in shortgrass steppe. In: Journal of Vegetation Science, DOI:10.1111/jvs.12508.

Clark, J., E. Grimm, J. Donovan, S. Fritz, D. Engrstom, and J. Almendinger. 2002. Drought cycles and landscape responses to past aridity on prairies of the Northern Great Plains, USA. Ecology, 83(3), Pages 595-601.

Connell, L. C., J. D. Scasta, and L. M. Porensky. 2018. Prairie dogs and wildfires shape vegetation structure in a sagebrush grassland more than does rest from ungulate grazing. Ecosphere 9(8):e02390. 10.1002/ecs2.2390

Collins, S. and S. Barber. (1985). Effects of disturbance on diversity in mixed-grass prairie. In: Vegetatio, 64, pages 87-94.

Egan, Timothy. 2006. The Worst Hard Time. Houghton Mifflin Harcourt Publishing Company, New York, NY.

Guyette, R.P., M.C. Stambaugh, D.C. Dey, and R.M. Muzika. 2012. Predicting fire frequency with chemistry and climate. In: Ecosystems, 15: pages 322-335.

Hart, R. and J. Hart. 1997. Rangelands of the Great Plains before European settlement. In: Rangelands, 19(1), pages 4-11.

Hart, R. 2001. Plant biodiversity on shortgrass steppe after 55 years of zero, light, moderate, or heavy cattle grazing. In: Plant Ecology, 155, pages 111-118.

Pellant, M., P. Shaver, D.A. Pyke, and J.E. Herrick. 2005. Interpreting indicators of rangeland health, Version 4. United States Department of the Interior, Bureau of Land Management.

Porensky, L.M. and D.M. Blumenthal. 2016. Historical wildfires do not promote cheatgrass invasion in a western Great Plains steppe. In: Biological Invasions 18:3333-3349: DOI 10.1007/s10530-16-1225-z

Porensky, L.M., J.D. Derner, and D.W. Pellatz. 2018. Plant community responses to historical wildfire in a shrubland-grassland ecotone reveal hybrid disturbance response. In: Ecosphere. DOI: 9(8):e02363. 10.1002/ecs2.2363.

Mack, Richard N., and J.N. Thompson. 1982. Evolution in steppe with few large, hooved mammals. In: The American Naturalist. 119, No. 6, pages 757-773.

Reyes-Fox, M., H. Stelzer, M.J. Trlica, G.S. McMaster, A.A. Andales, D.R. LeCain, and J.A. Morgan. 2014. Elevated CO2 further lengthens growing season under warming conditions. In: Nature, April 23, 2014. Available online. http://www.nature.com/nature/journal/v510/n7504/full/nature13207.html. Accessed 1 March, 2017.

Schoeneberger, P.J., D.A. Wysockie, E.C. Benham, and Soil Survey Staff. 2012. Field book for describing and sampling soils, Version 3.0. U.S. Dept. of Agriculture, Natural Resources Conservation Service.

Stahl, David W., E.R. Cook, M.K. Cleaveland, M.D. Therrell, D.M. Meko, H.D. Grissino-Mayer, E. Watson, and B.H. Luckman. Tree-ring data document 16th century megadrought over North America. 2000. In: Eos, 81(12), pages 121-125.

Stubbendieck, James, S.L. Hatch, and L.M. Landholt. 2003. North American wildland plants. Univ. of Nebraska Press, Lincoln and London.

Zelikova, T.J., D.M. Blumenthal, D.G. Williams, L. Souza, D.R. LeCain, and J. Morgan. 2014. Long-term exposure to elevated CO2 enhances plant community stability by suppressing dominant plant species in a mixed-grass prairie. In: Ecology, 2014. Available online. https://www.pnas.org/content/111/43/15456.

United States Department of Agriculture, Natural Resources Conservation Service. National Ecological Site Handbook, Title 190, Part 630, 1st Edition. Available online. https://directives.sc.egov.usda.gov/. Accessed 15 September, 2017.

United States Department of Agriculture, Natural Resources Conservation Service. 2009. Part 630, Hydrology, National Engineering Handbook.

United States Department of Agriculture, Natural Resources Conservation Service. 1972-2012. National Engineering Handbook Hydrology Chapters. http://www.nrcs.usda.gov/wps/portal/nrcs/detailfull/national/water/?&cid=stelprdb1043063 (Accessed August, 2015).

United States Department of Agriculture, Natural Resources Conservation Service. 1997, revised 2003.
National Range and Pasture Handbook. Available online. http://www.glti.nrcs.usda.gov/technical/publications/nrph.html
Accessed 26 February, 2018.

United States Department of Agriculture, Natural Resources Conservation Service. National Soil Survey Handbook title 430-VI. Available online. http://www.nrcs.usda.gov/wps/portal/nrcs/detail/soils/ref/?cid=nrcs142p2_054242.

United States Department of Agriculture, Natural Resources Conservation Service. Web Soil Survey. Available online. http://websoilsurvey.sc.egov.usda.gov/App/WebSoilSurvey.aspx. Accessed 15 November, 2017.

United States Department of Commerce, National Oceanic and Atmospheric Administration (NOAA). Cooperative climatological data summaries. NOAA Western Regional Climate Center, Reno, NV. Available online. http://www.wrcc.dri.edu/climatedata/climsum. Accessed 16 November, 2017.

Contributors

Everett Bainter
Glenn Mitchell

Approval

Kirt Walstad, 9/15/2024

Acknowledgments

Project Staff:
Kimberly Diller, Ecological Site Inventory Specialist, NRCS MLRA SSO, Pueblo CO
Mike Leno, Project Leader, NRCS MLRA SSO, Buffalo, WY

Partners/Contributors:
Joe Dyer, Soil Scientist, NRCS MLRA SSO, Buffalo, WY
Arnie Irwin, Soil Scientist, BLM, Buffalo, WY
Blaine Horn, Rangeland Extension Educator, UW Extension, Buffalo, WY
Isabelle Giuliani, Resource Soil Scientist, NRCS, Douglas, WY
Mary Jo Kimble, Project Leader, NRCS MLRA SSO, Miles City, MT
Ryan Murray, Rangeland Management Specialist, NRCS, Buffalo, WY
Lauren Porensky, Ph.D., Ecologist, ARS, Fort Collins, CO
Chadley Prosser, Rangeland Program Manager, USFS, Bismarck, ND
Bryan Christensen, Ecological Site Inventory Specialist, NRCS-MLRA SSO, Pinedale, WY
Marji Patz, Ecological Site Inventory Specialist, NRCS-MLRA SSO, Powell, WY
Rick Peterson, Ecological Site Inventory Specialist, NRCS-MLRA SSO, Rapid City, SD

Program Support:
John Hartung, WY State Rangeland Management Specialist-QC, NRCS, Casper, WY
David Kraft, NRCS MLRA Ecological Site Inventory Specialist-QA, Emporia, KS
Carla Green Adams, Editor, NRCS-SSR5, Denver, CO
Chad Remley, Regional Director, Northern Great Plains Soil Survey, Salina, KS

Rangeland health reference sheet

Interpreting Indicators of Rangeland Health is a qualitative assessment protocol used to determine ecosystem condition based on benchmark characteristics described in the Reference Sheet. A suite of 17 (or more) indicators are typically considered in an assessment. The ecological site(s) representative of an assessment location must be known prior to applying the protocol and must be verified based on soils and climate. Current plant community cannot be used to identify the ecological site.

Author(s)/participant(s)
Contact for lead author
Date	04/05/2005
Approved by	Kirt Walstad
Approval date
Composition (Indicators 10 and 12) based on	Annual Production