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Ecological site F115XC019IL

Clayey Floodplain Forest

Last updated: 12/30/2024
Accessed: 01/10/2025

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General information

Provisional. A provisional ecological site description has undergone quality control and quality assurance review. It contains a working state and transition model and enough information to identify the ecological site.

MLRA notes

Major Land Resource Area (MLRA): 115X–Central Mississippi Valley Wooded Slopes

This MLRA is characterized by deeply dissected, loess-covered hills bordering well defined valleys of the Illinois, Mississippi, Missouri, Ohio, and Wabash Rivers and their tributaries. It is used to produce cash crops and livestock. About one-third of the area is forested, mostly on the steeper slopes. This area is in Illinois (50 percent), Missouri (36 percent), Indiana (13 percent), and Iowa (1 percent) in two separate areas. It makes up about 25,084 square miles (64,967 square kilometers).

Most of this area is in the Till Plains section and the Dissected Till Plains section of the Central Lowland province of the Interior Plains. The Springfield-Salem plateaus section of the Ozarks Plateaus province of the Interior Highlands occurs along the Missouri River and the Mississippi River south of the confluence with the Missouri River. The nearly level to very steep uplands are dissected by both large and small tributaries of the Illinois, Mississippi, Missouri, Ohio, and Wabash Rivers. The Ohio River flows along the southernmost boundary of this area in Indiana. Well defined valleys with broad flood plains and numerous stream terraces are along the major streams and rivers. The flood plains along the smaller streams are narrow. Broad summits are nearly level to undulating. Karst topography is common in some parts along the Missouri and Mississippi Rivers and their tributaries. Well-developed karst areas have hundreds of sinkholes, caves, springs, and losing streams. In the St. Louis area, many of the karst features have been obliterated by urban development.

Elevation ranges from 90 feet (20 meters) on the southernmost flood plains to 1,030 feet (320 meters) on the highest ridges. Local relief is mainly 10 to 50 feet (3 to 15 meters) but can be 50 to 150 feet (15 to 45 meters) in the steep, deeply dissected hills bordering rivers and streams. The bluffs along the major rivers are generally 200 to 350 feet (60 to 105 meters) above the valley floor.

The uplands in this MLRA are covered almost entirely with Peoria Loess. The loess can be more than 7 feet (2 meters) thick on stable summits. On the steeper slopes, it is thin or does not occur. In Illinois, the loess is underlain mostly by Illinoian-age till that commonly contains a paleosol. Pre-Illinoian-age till is in parts of this MLRA in Iowa and Missouri and to a minor extent in the western part of Illinois. Wisconsin-age outwash, alluvial deposits, and sandy eolian material are on some of the stream terraces and on dunes along the major tributaries. The loess and glacial deposits are underlain by several bedrock systems. Pennsylvanian and Mississippian bedrock are the most extensive. To a lesser extent are Silurian, Devonian, Cretaceous, and Ordovician bedrock. Karst areas have formed where limestone is near the surface, mostly in the southern part of the MLRA along the Mississippi River and some of its major tributaries. Bedrock outcrops are common on the bluffs along the Mississippi, Ohio, and Wabash Rivers and their major tributaries and at the base of some steep slopes along minor streams and drainageways.

The annual precipitation ranges from 35 to 49 inches (880 to 1,250 millimeters) with a mean of 41 inches (1,050 millimeters). The annual temperature ranges from 48 to 58 degrees F (8.6 to 14.3 degrees C) with a mean of 54 degrees F (12.3 degrees C). The freeze-free period ranges from 150 to 220 days with a mean of 195 days.

Soils The dominant soil orders are Alfisols and, to a lesser extent, Entisols and Mollisols. The soils in the area have a mesic soil temperature regime, an aquic or udic soil moisture regime, and mixed or smectitic mineralogy. They are shallow to very deep, excessively drained to poorly drained, and loamy, silty, or clayey.

The soils on uplands in this area support natural hardwoods. Oak, hickory, and sugar maple are the dominant species. Big bluestem, little bluestem, and scattered oak and eastern redcedar grow on some sites. The soils on flood plains support mixed forest vegetation, mainly American elm, eastern cottonwood, river birch, green ash, silver maple, sweetgum, American sycamore, pin oak, pecan, and willow. Sedge and grass meadows and scattered trees are on some low-lying sites. (United States Department of Agriculture, Natural Resources Conservation Service, 2022)

LRU notes

The Central Mississippi Valley Wooded Slopes, Northern part (Land Resource Unit (LRU) (115XC) encompasses the Wyaconda River Dissected Till Plains, Mississippi River Hills, and Mississippi River Alluvial Plain (Schwegman et al. 1973; Nelson 2010). It spans three states – Illinois (73 percent), Iowa (6 percent), and Missouri (21 percent) – comprising about 13,650 square miles (Figure 1). The elevation ranges from 420 feet above sea level (ASL) along the Mississippi River floodplains to 885 feet on the upland ridges. Local relief varies from 10 to 20 feet but can be as high as 50 to 100 feet along drainageways and streams and the bluffs on the major rivers reaching 250 feet above valley floors. Wisconsin-aged loess covers the uplands, while Illinoian glacial drift lies directly below. The loess and drift deposits are underlain by several bedrock systems, including the Cretaceous, Pennsylvania, Mississippian, Silurian, Devonian, and Ordovician Systems. Wisconsin outwash deposits and sandy eolian material occur along stream terraces of major tributaries (USDA-NRCS 2006).

The vegetation across the region has undergone drastic changes over time. At the end of the last glacial episode – the Wisconsinan glaciation – the evolution of vegetation began with the development of tundra habitats, followed by a phase of spruce and fir forests, and eventually spruce-pine forests. Not until approximately 9,000 years ago did the climate undergo a warming trend which prompted the development of deciduous forests dominated by oak and hickory. As the climate continued to warm and dry, prairies began to develop approximately 8,300 years ago. Another shift in climate that resulted in an increase in moisture prompted the emergence of savanna-like habitats from 8,000 to 5,000 years before present (Taft et al. 2009). During the most recent climatic shifts, forested ecosystems maintained footholds on steep valley sides and wet floodplains. Due to the physiography of the MLRA, forests were the dominant ecosystems and were affected by such natural disturbances as droughts, wind, lightning, and occasional fire (Taft et al. 2009).

Classification relationships

USFS Subregions: Central Dissected Till Plains (251C)Section; Western Mississippi River Hills (251Ce), Mississippi River and Illinois Alluvial Plains (251Cf), Eastern Mississippi River Hills (251Ci), Galesburg Dissected Till Plain (251Cj), and Wyaconda River Dissected Till Plain (251Cm) Subsections (Cleland et al. 2007)

U.S. EPA Level IV Ecoregion: Upper Mississippi River Alluvial Plain (72d), River Hills (72f), and Western Dissected Illinoian Till Plain (72i) (USEPA 2013)

National Vegetation Classification - Plant Associations: Acer saccharinum – Fraxinus pensylvanica – Ulmus americana Floodplain Forest (CEGL002586) (Nature Serve 2015)

Biophysical Settings: Central Interior and Appalachian Floodplain Systems (BpS 4214710) (LANDFIRE 2009)

Natural Resources Conservation Service – Iowa Plant Community Species List: Forest, Silver Maple - Elm (USDA-NRCS 2007)

Illinois Natural Areas Inventory: Wet Floodplain Forest (White and Madany 1978)

Iowa Department of Natural Resources: Floodplain Forest (INAI 1984)

Missouri Terrestrial Natural Communities: Wet bottomland forest (Nelson 2010)

Ecological site concept

Clayey Floodplain Forests occur on floodplains in river valleys. The soils are Mollisols that are poorly drained and very deep, formed in clayey alluvium.

The historic pre-European settlement vegetation on this ecological site was dominated by deciduous trees and a sparse understory of shade-tolerant herbaceous plants. The tree canopy is comprised of American sycamore (Platanus occidentalis L.) and silver maple (Acer saccharinum L.). Other tree species that may occur include green ash (Fraxinus pennsylvanica Marshall), bur oak (Quercus macrocarpa Michx.), American elm (Ulmus americana L), and boxelder (Acer negundo L.). Eastern poison ivy (Toxicodendron radicans L.) is a common climbing vine. The understory is mostly bare ground and leaf litter, but few herbaceous species are present. Rice cutgrass (Leersia oryzoides (L.) Sw.) may be a frequently encountered herbaceous species. Flooding is the primary disturbance factor that maintains this site, while damage from storms is a secondary disturbance (LANDFIRE 2009).

Associated sites

F115XC021IL	Sandy Floodplain Forest Sandy and gravelly alluvial parent material including Caneek variant, Elsah, Fruitfield, Klum, Landes, Psamments, Sarpy, and Zumbro soils
F115XC020IL	Loamy Floodplain Forest Silty and loamy alluvial parent material including Ackmore, Ambraw, Arenzville, Belknap, Blake, Blyton, Ceresco, Coffeen, Dockery, Dorchester, Dozaville, Elrick, Haymond, Huntsville, Jules, Lawson, Medway, Orion, Paxico, Radford, Riley, Ross, Shaffton, Tice, Udorthents, Volney, Wakeland, and Wirt soils

F115XC021IL

Sandy Floodplain Forest

Sandy and gravelly alluvial parent material including Caneek variant, Elsah, Fruitfield, Klum, Landes, Psamments, Sarpy, and Zumbro soils

F115XC020IL

Loamy Floodplain Forest

Silty and loamy alluvial parent material including Ackmore, Ambraw, Arenzville, Belknap, Blake, Blyton, Ceresco, Coffeen, Dockery, Dorchester, Dozaville, Elrick, Haymond, Huntsville, Jules, Lawson, Medway, Orion, Paxico, Radford, Riley, Ross, Shaffton, Tice, Udorthents, Volney, Wakeland, and Wirt soils

Similar sites

F115XC021IL	Sandy Floodplain Forest Sandy Floodplain Forests are in similar landscape positions but parent material is sandy and gravelly alluvium
F115XC020IL	Loamy Floodplain Forest Loamy Floodplain Forests are in similar landscape positions but parent material is silty and loamy alluvium

F115XC021IL

Sandy Floodplain Forest

Sandy Floodplain Forests are in similar landscape positions but parent material is sandy and gravelly alluvium

F115XC020IL

Loamy Floodplain Forest

Loamy Floodplain Forests are in similar landscape positions but parent material is silty and loamy alluvium

Table 1. Dominant plant species

Tree	(1) Platanus occidentalis (2) Acer saccharinum
Shrub	(1) Toxicodendron radicans
Herbaceous	(1) Leersia oryzoides

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Physiographic features

Clayey Floodplain Forests occur on floodplains in river valleys. They are situated on elevations ranging from approximately 341 to 1000 feet ASL. The site experiences occasional flooding that can last up to 7 days.

Figure 1. Location of Clayey Floodplain Forest ecological site within LRU 115XC.

Figure 2. Representative block diagram of Clayey Floodplain Forest and associated ecological sites.

Table 2. Representative physiographic features

Slope shape across	(1) Linear
Slope shape up-down	(1) Linear
Landforms	(1) River valley > Flood plain
Runoff class	Low to very high
Flooding duration	Brief (2 to 7 days)
Flooding frequency	Occasional
Ponding frequency	None
Elevation	341 – 1,000 ft
Slope	2%
Water table depth	6 in
Aspect	Aspect is not a significant factor

Climatic features

The Central Mississippi Valley Wooded Slopes, Northern Part falls into the humid subtropical (Cfa) and hot-summer humid continental climate (Dfa) Köppen-Geiger climate classifications (Peel et al. 2007). The two main factors that drive the climate of the MLRA are latitude and weather systems. Latitude, and the subsequent reflection of solar input, determines air temperatures and seasonal variations. Solar energy varies across the seasons, with summer receiving three to four times as much energy as opposed to winter. Weather systems (air masses and cyclonic storms) are responsible for daily fluctuations of weather conditions. High-pressure systems are responsible for settled weather patterns where sun and clear skies dominate. In fall, winter, and spring, the polar jet stream is responsible for the creation and movement of low-pressure systems. The clouds, winds, and precipitation associated with a low-pressure system regularly follow high-pressure systems every few days (Angel n.d.).

The soil temperature regime of LRU 115XC is classified as mesic, where the mean annual soil temperature is between 46 and 59°F (USDA-NRCS 2006). Temperature and precipitation occur along a north-south gradient, where temperature and precipitation increase the further south one travels. The average freeze-free period of this ecological site is about 184 days, while the frost-free period is about 152 days. The majority of the precipitation occurs as rainfall in the form of convective thunderstorms during the growing season. Average annual precipitation is 38 inches, which includes rainfall plus the water equivalent from snowfall. The average annual low and high temperatures are 42 and 62°F, respectively.

Table 3. Representative climatic features

Frost-free period (characteristic range)	141-168 days
Freeze-free period (characteristic range)	170-196 days
Precipitation total (characteristic range)	35-40 in
Frost-free period (actual range)	140-170 days
Freeze-free period (actual range)	169-203 days
Precipitation total (actual range)	35-40 in
Frost-free period (average)	152 days
Freeze-free period (average)	184 days
Precipitation total (average)	38 in

Characteristic range

Actual range

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Figure 3. Monthly precipitation range

Characteristic range

Actual range

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Figure 4. Monthly minimum temperature range

Characteristic range

Actual range

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Figure 5. Monthly maximum temperature range

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Figure 6. Monthly average minimum and maximum temperature

Figure 7. Annual precipitation pattern

Figure 8. Annual average temperature pattern

Climate stations used

(1) CLINTON #1 [USC00131635], Camanche, IA
(2) NEW BOSTON DAM 17 [USC00116080], Wapello, IL
(3) CANTON L&D 20 [USC00231275], Canton, MO
(4) CLARKSVILLE L&D 24 [USC00231640], Clarksville, MO
(5) HAVANA [USC00113940], Lewistown, IL

Influencing water features

Clayey Floodplain Forests are classified as a RIVERINE: Frequently Flooded; forested wetland under the Hydrogeomorphic (HGM) classification system (Smith et al. 1995; USDA-NRCS 2008) and as a Palustrine, Forested, Broad-leaved Deciduous, Temporarily Flooded wetland under the National Wetlands Inventory (FGDC 2013). Overbank flow from the channel and subsurface hydraulic connections are the main sources of water for this ecological site (Smith et al. 1995). Infiltration is very slow (Hydrologic Group D) for undrained soils, and surface runoff is low to very high.

Wetland description

Primary wetland hydrology indicators for an intact Clayey Floodplain Forest may include: A1 Surface water, A2 High water table, A3 Saturation, B1 Water marks, B2 Sediment deposits, B3 Drift deposits, B8 Sparsely vegetated concave surface, and B9 Water-stained leaves. Secondary wetland hydrology indicators may include: D5 FAC-neutral test (USACE 2010).

Figure 9. Hydrologic cycling in Clayey Floodplain Forest ecological site.

Soil features

Soils of Clayey Floodplain Forests are in the Mollisols order, further classified as Fluvaquentic Endoaquolls and Vertic Endoaquolls with very slow infiltration and low to very high runoff potential. The soil series associated with this site includes Carlow, Chequest, McFain, Moline, and Titus. The parent material is clayey alluvium, and the soils are poorly drained and very deep with seasonal high-water tables. Soil pH classes are strongly acid to moderately alkaline. No rooting restrictions are noted for the soils of this ecological site.

Some soil map units in this ecological site, if not drained, may meet the definition of hydric soils and are listed as meeting criteria 2 of the hydric soils list (77 FR 12234).

Figure 10. Profile sketches of soil series associated with Clayey Floodplain Forest.

Table 4. Representative soil features

Parent material	(1) Alluvium
Surface texture	(1) Silty clay (2) Silty clay loam
Family particle size	(1) Fine (2) Clayey over loamy
Drainage class	Poorly drained
Permeability class	Very slow to slow
Depth to restrictive layer	80 in
Soil depth	80 in
Surface fragment cover <=3"	Not specified
Surface fragment cover >3"	Not specified
Available water capacity (Depth not specified)	4 – 8 in
Calcium carbonate equivalent (Depth not specified)	40%
Electrical conductivity (Depth not specified)	2 mmhos/cm
Sodium adsorption ratio (Depth not specified)	Not specified
Soil reaction (1:1 water) (Depth not specified)	5.1 – 8.4
Subsurface fragment volume <=3" (Depth not specified)	3%
Subsurface fragment volume >3" (Depth not specified)	Not specified

Ecological dynamics

The information in this Ecological Site Description, including the state-and-transition model (STM), was developed based on historical data, current field data, professional experience, and a review of the scientific literature. As a result, all possible scenarios or plant species may not be included. Key indicator plant species, disturbances, and ecological processes are described to inform land management decisions.

The MLRA lies within the tallgrass prairie ecosystem of the Midwest, but a variety of environmental and edaphic factors resulted in a landscape that historically supported upland hardwood forests, lowland mixed forests, and scattered grass and sedge meadows. Clayey Floodplain Forests form an aspect of this vegetative continuum. This ecological site occurs on floodplains on poorly drained alluvial soils. Species characteristic of this ecological site consist of hydrophytic woody and herbaceous vegetation.

Flooding is the dominant disturbance factor in Clayey Floodplain Forests, and storm damage is a secondary disturbance. Periodic flooding occurs seasonally, sometimes lasting up to a month at a time. Damage to trees from wind storms can vary from minor, patchy effects of individual trees to stand effects that temporarily affect community structure and species richness and diversity (Irland 2000; Peterson 2000).

Today, many Clayey Floodplain Forests have been reduced as a result of conversion to pasture. A few sites have been cleared and drained for agricultural production. Remnant sites have been degraded due to significant changes to the natural hydrologic regime and diminished water quality in the watershed. The state-and-transition model that follows provides a detailed description of each state, community phase, pathway, and transition. This model is based on available experimental research, field observations, literature reviews, professional consensus, and interpretations.

State and transition model

More interactive model formats are also available. View Interactive Models
Click on state and transition labels to scroll to the respective text

Ecosystem states

1. Reference State

2. Hydrologically-altered State

3. Pasture State

4. Cropland State

5. Reconstructed Floodplain Forest State

States 2 and 5 (additional transitions)

2. Hydrologically-altered State

5. Reconstructed Floodplain Forest State

T1A	-	Changes to natural hydroperiod and/or land abandonment
T1B	-	Cultural treatments are implemented to increase forage quality and yield
T1C	-	Agricultural conversion via tillage, seeding, and non-selective herbicide
T2A	-	Cultural treatments are implemented to increase forage quality and yield
T2B	-	Agricultural conversion via tillage, seeding, and non-selective herbicide
R2A	-	Site preparation, tree planting, repair hydrology, non-native species control
T3A	-	Changes to natural hydroperiod and/or land abandonment
T3B	-	Agricultural conversion via tillage, seeding, and non-selective herbicide
R3A	-	Site preparation, tree planting, repair hydrology, non-native species control
T4A	-	Changes to natural hydroperiod and/or land abandonment
T4B	-	Cultural treatments are implemented to increase forage quality and yield
R4A	-	Site preparation, tree planting, repair hydrology, non-native species control
T5A	-	Changes to natural hydroperiod and/or land abandonment
T5B	-	Cultural treatments are implemented to increase forage quality and yield
T5C	-	Agricultural conversion via tillage, seeding, and non-selective herbicide

State 1 submodel, plant communities

1.1. American Sycamore – Silver Maple/Eastern Poison Ivy/Rice Cutgrass – bare ground

1.2. American Sycamore – Bur Oak/Common Hackberry/Rice Cutgrass – Canadian Woodnettle

1.1A	-	Major Flood Event
1.2A	-	Natural succession as a result of no disturbances

State 2 submodel, plant communities

2.1. Silver Maple – American Elm/Green Ash – Common Hackberry/Canadian Woodnettle – Creeping Jenny

State 3 submodel, plant communities

3.1. Hayfield

3.2. Smooth brome -Tall fescue - Clover Pasture

3.3. Broomsedge -Thistle-Smooth brome -Tall fescue Pasture

3.1A	-	Grazing; proper animal to forage balance.
3.1B	-	Grazing; Overutilization of forage plants.
3.2A	-	Mechanical harvesting
3.2B	-	Grazing; Overutilization of the forage plant

State 4 submodel, plant communities

4.1. Conventional Tillage Field

4.2. Conservation Tillage Field

4.3. Conservation Tillage Field/ Cover Crop Field

4.1A	-	Less tillage, residue management
4.1B	-	Less tillage, residue management, and implementation of cover cropping
4.2A	-	Intensive tillage, remove residue, and reinitiate monoculture row cropping
4.2B	-	Implementation of cover cropping
4.3B	-	Intensive tillage, remove residue, and reinitiate monoculture row cropping
4.3A	-	Remove cover cropping

State 5 submodel, plant communities

5.1. Early Successional Reconstructed Floodplain Forest

5.2. Late Successional Reconstructed Floodplain Forest

5.1A	-	Timber stand improvement practices implemented
5.2A	-	Restoration setback

State 1
Reference State

The reference plant community is categorized as a floodplain forest community, dominated by hydrophytic woody and herbaceous vegetation. The two community phases within the reference state are dependent on periodic flooding. The amount and duration of floodwaters alters species composition, cover, and extent. Damage from storms have more localized impacts in the reference phases, but does contribute to overall species composition, diversity, cover, and productivity.

Dominant plant species

American sycamore (Platanus occidentalis), tree
silver maple (Acer saccharinum), tree
eastern poison ivy (Toxicodendron radicans), shrub
Bare Ground (Bare Ground), other herbaceous
rice cutgrass (Leersia oryzoides), other herbaceous

Community 1.1
American Sycamore – Silver Maple/Eastern Poison Ivy/Rice Cutgrass – bare ground

Sites in this reference community phase are a closed canopy forest (80 to 100 percent cover) dominated by American sycamore and silver maple, with subdominants including American elm, green ash, bur oak, and boxelder. Trees are large (21 to 33-inch DBH) and range in height from 30 to over 80 feet tall (LANDFIRE 2009). Climbing vines can be present, typically including eastern poison ivy. The herbaceous layer is sparse and lacking in species diversity. Rice cutgrass may be the dominant species present, but bare ground and litter comprise the majority of the ground cover. Periodic flooding will maintain this phase, but over time the site may accumulate more sediments causing it to become more elevated and thereby shifting the community to phase 1.2.

Dominant plant species

American sycamore (Platanus occidentalis), tree
silver maple (Acer saccharinum), tree
eastern poison ivy (Toxicodendron radicans), shrub
rice cutgrass (Leersia oryzoides), grass
Bare Ground (Bare Ground), other herbaceous

Community 1.2
American Sycamore – Bur Oak/Common Hackberry/Rice Cutgrass – Canadian Woodnettle

This reference community phase represents the plant community following long-term sediment accumulation and site elevation. American sycamore is still present, but bur oak and American elm become co-dominant. Common hackberry can be a common canopy and sub-canopy component. The elevation increase on the site reduces the frequency of flooding and allows the herbaceous layer to become more continuous with species such as rice cutgrass, eastern waterleaf (Hydrophyllum virginianum L.), Canadian clearweed (Pilea pumila (L.) A. Gray), and Canadian woodnettle (Laportea canadensis (L.) Weddell) (White and Madany 1978).

Dominant plant species

American sycamore (Platanus occidentalis), tree
bur oak (Quercus macrocarpa), tree
common hackberry (Celtis occidentalis), shrub
rice cutgrass (Leersia oryzoides), grass
Canadian woodnettle (Laportea canadensis), other herbaceous

Pathway 1.1A
Community 1.1 to 1.2

Natural succession from sediment accumulation.

Pathway 1.2A
Community 1.2 to 1.1

Major disturbance event such as flood or windthrow.

State 2
Hydrologically-altered State

Agricultural tile drainage, stream channelization, and levee construction in hydrologically-connected waters have drastically changed the natural hydrologic regime of Floodplain Forests. In addition, increased amounts of precipitation and intensity have amplified flooding events (Pryor et al. 2014). This has resulted in a type conversion from the species-rich forest to a ruderal floodplain forest state. In addition, exotic species have encroached and continuously spread, reducing native diversity and ecosystem stability (Eggers and Reed 2015).

Dominant plant species

silver maple (Acer saccharinum), tree
American elm (Ulmus americana), tree
green ash (Fraxinus pennsylvanica), tree
common hackberry (Celtis occidentalis), tree
Canadian woodnettle (Laportea canadensis), other herbaceous
creeping jenny (Lysimachia nummularia), other herbaceous

Community 2.1
Silver Maple – American Elm/Green Ash – Common Hackberry/Canadian Woodnettle – Creeping Jenny

This community phase represents a transition in plant community composition as a result of an altered hydrologic regime. Silver maple and American elm are the dominant canopy species, and green ash, common hackberry, and boxelder are common canopy and subcanopy associates. The herbaceous layer may be nearly continuous but lacks in species diversity. Disturbance-tolerant natives, such as Canadian woodnettle, can be accompanied by aggressive non-native invaders, such as creeping jenny (Lysimachia nummularia L.) and garlic mustard (Alliaria petiolata (M. Bieb.) Cavara & Grande).

Dominant plant species

silver maple (Acer saccharinum), tree
American elm (Ulmus americana), tree
green ash (Fraxinus pennsylvanica), tree
common hackberry (Celtis occidentalis), tree
Canadian woodnettle (Laportea canadensis), other herbaceous
creeping jenny (Lysimachia nummularia), other herbaceous

State 3
Pasture State

The pasture state arises when the site is converted to a farming system that emphasizes domestic livestock production, known as grassland agriculture. Fire suppression, periodic cultural treatments (e.g., clipping, drainage, soil amendment applications, planting new species and/or cultivars, mechanical harvesting) and grazing by domesticated livestock transition and maintain this state (USDA-NRCS 2003). Early settlers seeded non-native species, as smooth brome (Bromus inermis Leyss.), tall fescue ( Festuca arundinacea) and Kentucky bluegrass (Poa pratensis L.), to help extend the grazing season (Smith 1998). Over time, as lands were continuously harvested or grazed by herds of cattle, these species were able to spread and expand across the landscape, reducing the native species diversity and ecological function.

Dominant plant species

smooth brome (Bromus inermis), grass
tall fescue (Schedonorus arundinaceus), grass
Kentucky bluegrass (Poa pratensis), grass
red clover (Trifolium pratense), other herbaceous
white clover (Trifolium repens), other herbaceous

Community 3.1
Hayfield

Sites in this community phase consist of forage plants that are planted and mechanically harvested. Mechanical harvesting removes a significant portion of the aboveground biomass and nutrients that feed the soil microorganisms (Franzluebbers et al. 2000; USDA-NRCS 2003). As a result, soil biology is reduced leading to decreases in nutrient uptake by plants, soil organic matter, and soil aggregation. Frequent biomass removal can also reduce the site’s carbon sequestration capacity (Skinner 2008). Many species can be seeded and the plant selection will depend on the landowner objectives.

Dominant plant species

smooth brome (Bromus inermis), grass
tall fescue (Schedonorus arundinaceus), grass
Kentucky bluegrass (Poa pratensis), grass
timothy (Phleum pratense), grass
orchardgrass (Dactylis glomerata), grass
alfalfa (Medicago), other herbaceous
clover (Trifolium), other herbaceous

Community 3.2
Smooth brome -Tall fescue - Clover Pasture

This community is characterized by seeded cool-season grass and forbs. Species will depend upon landowner goals and objectives and may include many different grasses and forbs. Common species include smooth brome (Bromus inermis), tall fescue (Festuca arundinacea), Kentucky bluegrass (Poa pratensis), timothy (Phleum pratense), red clover ( Trifolium pratense) and white clover (Trifolium repens L.). Management inputs include control of weeds and brush. These sites are managed to ensure a proper forage/animal balance. Plants are not overutilized and have adequate rest and recovery.

Dominant plant species

tall fescue (Schedonorus arundinaceus), grass
smooth brome (Bromus inermis), grass
Kentucky bluegrass (Poa pratensis), grass
white clover (Trifolium repens), other herbaceous
red clover (Trifolium pratense), other herbaceous

Community 3.3
Broomsedge -Thistle-Smooth brome -Tall fescue Pasture

Over utilization of the pasture will result in a shift to include more undesirable species such as thistle (Cirsium spp.), broomsedge (Andropogon virginicus L.), ironweed (Vernonia gigantea), buttercup (Ranunculus spp.), ragweed (Ambrosia spp.) and blackberries (Rubus spp.). Many woody and weed species may be present depending on seed sources and level of soil disturbance. This community reflects an improper forage-to-animal balance which will negatively impact forage productivity and reproduction, soil health, and water quality. Ecological resiliency is compromised under these conditions.

Dominant plant species

broomsedge bluestem (Andropogon virginicus), grass
sedge (Carex), grass
tall fescue (Schedonorus arundinaceus), grass
smooth brome (Bromus inermis), grass
thistle (Cirsium), other herbaceous
buttercup (Ranunculus), other herbaceous
ironweed (Vernonia), other herbaceous
ragweed (Ambrosia), other herbaceous

Pathway 3.1A
Community 3.1 to 3.2

Mechanical harvesting is replaced with domestic livestock grazing.

Pathway 3.1B
Community 3.1 to 3.3

Mechanical harvesting is replaced with domestic livestock grazing. Overutilization of the forage plants.

Pathway 3.2A
Community 3.2 to 3.1

Domestic livestock are removed and mechanical harvesting is implemented.

Pathway 3.2B
Community 3.2 to 3.3

Grazing of livestock with overutilization of the forage plant

Pathway 3.3B
Community 3.3 to 3.1

Domestic livestock are removed and mechanical harvesting is implemented.

Pathway 3.3A
Community 3.3 to 3.2

Forage plants are not overutilized and the site has a proper forage-to-animal balance.

State 4
Cropland State

The Midwest is well-known for its highly-productive agricultural soils, and as a result, much of the MLRA has been converted to cropland, including portions of this ecological site. The continuous use of tillage, row-crop planting, and chemicals (i.e., herbicides, fertilizers, etc.) has effectively eliminated the reference community and many of its natural ecological functions in favor of crop production. Corn (Zea mays L.) and soybeans (Glycine max (L.) Merr.) are the dominant crops for the site. These areas are likely to remain in crop production for the foreseeable future.

Dominant plant species

corn (Zea mays), other herbaceous
soybean (Glycine max), other herbaceous

Community 4.1
Conventional Tillage Field

Sites in this community phase typically consist of monoculture row-cropping maintained by conventional tillage practices. They are cropped in either continuous corn or corn-soybean rotations. The frequent use of deep tillage, low crop diversity, and bare soil conditions during the non-growing season negatively impacts soil health. Under these practices, soil aggregation is reduced or destroyed, soil organic matter is reduced, erosion and runoff are increased, and infiltration is decreased, which can ultimately lead to undesirable changes in the hydrology of the watershed (Tomer et al. 2005).

Dominant plant species

corn (Zea mays), other herbaceous
soybean (Glycine max), other herbaceous

Community 4.2
Conservation Tillage Field

This community phase is characterized by rotational crop production that utilizes various conservation tillage methods to promote soil health and reduce erosion. Conservation tillage methods include strip-till, ridge-till, vertical-till, or no-till planting systems. Strip-till keeps seedbed preparation to narrow bands less than one-third the width of the row where crop residue and soil consolidation are left undisturbed in-between seedbed areas. Strip-till planting may be completed in the fall and nutrient application either occurs simultaneously or at the time of planting. Ridge-till uses specialized equipment to create ridges in the seedbed and vegetative residue is left on the surface in between the ridges. Weeds are controlled with herbicides and/or cultivation, seedbed ridges are rebuilt during cultivation, and soils are left undisturbed from harvest to planting. Vertical-till systems employ machinery that lightly tills the soil and cuts up crop residue, mixing some of the residue into the top few inches of the soil while leaving a large portion on the surface. No-till management is the most conservative, disturbing soils only at the time of planting and fertilizer application. Compared to conventional tillage systems, conservation tillage methods can reduce soil erosion, increase organic matter and water availability, improve water quality, and reduce soil compaction.

Dominant plant species

corn (Zea mays), other herbaceous
soybean (Glycine max), other herbaceous

Community 4.3
Conservation Tillage Field/ Cover Crop Field

This condition applies conservation tillage methods as described above as well as adds cover crop practices. Cover crops typically include nitrogen-fixing species (e.g., legumes), small grains (e.g., rye, wheat, oats), or forage covers (e.g., turnips, radishes, rapeseed). The addition of cover crops not only adds plant diversity but also promotes soil health by reducing soil erosion, limiting nitrogen leaching, suppressing weeds, increasing soil organic matter, and improving the overall soil. In the case of small grain cover crops, surface cover and water infiltration are increased, while forage covers can be used to graze livestock or support local wildlife. Of the three community phases for this state, this phase promotes the greatest soil sustainability and improves ecological functioning within a cropland system.

Dominant plant species

oat (Avena), grass
wheat (Triticum), grass
rye (Secale), grass
corn (Zea mays), other herbaceous
soybean (Glycine max), other herbaceous
radish (Raphanus), other herbaceous

Pathway 4.1A
Community 4.1 to 4.2

Tillage operations are greatly reduced, crop rotation occurs on a regular schedule, and crop residue is allowed to remain on the soil surface.

Pathway 4.1B
Community 4.1 to 4.3

Tillage operations are greatly reduced or eliminated, crop rotation is either reduced or eliminated, and crop residue is allowed to remain on the soil surface, and cover crops are implemented to prevent soil erosion.

Pathway 4.2A
Community 4.2 to 4.1

Intensive tillage is utilized and monoculture row-cropping is established.

Pathway 4.2B
Community 4.2 to 4.3

Cover crops are implemented to prevent soil erosion.

Pathway 4.3B
Community 4.3 to 4.1

Intensive tillage is utilized, cover crops practices are abandoned, monoculture row-cropping is established, and crop rotation is reduced or eliminated.

Pathway 4.3A
Community 4.3 to 4.2

Cover crop practices are abandoned.

State 5
Reconstructed Floodplain Forest State

The combination of natural and anthropogenic disturbances occurring today has resulted in numerous ecosystem health issues, and restoration back to the historic reference state may not be possible. Many natural forest communities are being stressed by non-native diseases and pests, habitat fragmentation, permanent changes in hydrologic regimes, and overabundant deer populations on top of naturally-occurring disturbances (severe weather and native pests) (Flickinger 2010). However, these habitats provide multiple ecosystem services including carbon sequestration; clean air and water; soil conservation; biodiversity support; wildlife habitat; as well as a variety of cultural activities (e.g., hiking, hunting) (Millennium Ecosystem Assessment 2005; Flickinger 2010). Therefore, conservation of floodplain forests should still be pursued. Habitat reconstructions are an important tool for repairing natural ecological functioning and providing habitat protection for numerous species of Clayey Floodplain Forests. Therefore, ecological restoration should aim to aid the recovery of degraded, damaged, or destroyed ecosystems. A successful restoration will have the ability to structurally and functionally sustain itself, demonstrate resilience to the ranges of stress and disturbance, and create and maintain positive biotic and abiotic interactions (SER 2002). The reconstructed forest state is the result of a long-term commitment involving a multi-step, adaptive management process.

Dominant plant species

maple (Acer), tree
American sycamore (Platanus occidentalis), tree

Community 5.1
Early Successional Reconstructed Floodplain Forest

This community phase represents the early community assembly from forest reconstruction. It is highly dependent on the current condition of the site based on past and current land management actions, invasive species, and proximity to land populated with non-native pests and diseases. Therefore, no two sites will have the same early successional composition. Technical forestry assistance should be sought to develop suitable conservation management plans.

Dominant plant species

maple (Acer), tree
American sycamore (Platanus occidentalis), tree

Community 5.2
Late Successional Reconstructed Floodplain Forest

Appropriately timed management practices (e.g. forest stand improvement, continuing integrated pest management) applied to the early successional community phase can help increase the stand maturity, pushing the site into a late successional community phase over time. A late successional reconstructed forest will have an uneven-aged, closed canopy and a well-developed understory.

Dominant plant species

maple (Acer), tree
American sycamore (Platanus occidentalis), tree

Pathway 5.1A
Community 5.1 to 5.2

Application of stand improvement practices in line with a developed management plan.

Pathway 5.2A
Community 5.2 to 5.1

Reconstruction experiences a setback from extreme weather event or improper timing of management actions.

Transition T1A
State 1 to 2

Transition 1A – Altered hydrology throughout the watershed transitions the site to the hydrologically-altered state (2).

Transition T1B
State 1 to 3

Transition 1B – Woody species removal and cultural treatments to enhance forage quality and yield transition the site to the pasture state (3).

Transition T1C
State 1 to 4

Transition 1C – Woody species removal, tillage, seeding of agricultural crops, and non-selective herbicide transition the site to the cropland state (4).

Transition T2A
State 2 to 3

Transition 2A – Woody species removal and cultural treatments to enhance forage quality and yield transition the site to the pasture state (3).

Transition T2B
State 2 to 4

Transition 2B – Woody species removal, tillage, seeding of agricultural crops, and non-selective herbicide transition the site to the cropland state (4).

Restoration pathway R2A
State 2 to 5

Restoration 2A – Site preparation, tree planting, timber stand improvement, non-native species control, and water control structures installed to improve and regulate hydrology transition this site to the reconstructed forest state (5).

Transition T3A
State 3 to 2

Transition 3A – Land is abandoned and left fallow; natural succession by opportunistic species transition this site the hydrologically-altered state (2).

Transition T3B
State 3 to 4

Transition 3B – Tillage, seeding of agricultural crops, and non-selective herbicide transition the site to the cropland state (4).

Restoration pathway R3A
State 3 to 5

Restoration 3A – Site preparation, tree planting, timber stand improvement, non-native species control, and water control structures installed to improve and regulate hydrology transition this site to the reconstructed forest state (5).

Transition T4A
State 4 to 2

Transition 4A – Land abandonment transitions the site to the hydrologically-altered state (2).

Transition T4B
State 4 to 3

Transition 4B – Cultural treatments to enhance forage quality and yield transition the site to the pasture state (3).

Restoration pathway R4A
State 4 to 5

Restoration 4A – Site preparation, tree planting, timber stand improvement, non-native species control, and water control structures installed to improve and regulate hydrology transition this site to the reconstructed forest state (5).

Restoration pathway T5A
State 5 to 2

Transition 5A – Removal of water control structures and unmanaged invasive species populations transition this site to the hydrologically-altered state (2).

Transition T5B
State 5 to 3

Transition 5B – Tree removal and cultural treatments to enhance forage quality and yield transition the site to the pasture state (3).

Transition T5C
State 5 to 4

Transition 5C – Tree removal, tillage, seeding of agricultural crops, and non-selective herbicide transition this site to the cropland state (4).

Additional community tables

Supporting information

Inventory data references

No field plots were available for this site. A review of the scientific literature and professional experience were used to approximate the plant communities for this provisional ecological site. Information for the state-and-transition model was obtained from the same sources. All community phases are considered provisional based on these plots and the sources identified in this ecological site description.

Other references

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Bharati, L., K.-H. Lee, T.M. Isenhart, and R.C. Schultz. 2002. Soil-water infiltration under crops, pasture, and established riparian buffer in Midwestern USA. Agroforestry Systems 56: 249-257.

Changes in Hydric Soils Database Selection Criteria. 77 Federal Register 12234 (29 February 2012), pp. 12234-12235.

Cleland, D.T., J.A. Freeouf, J.E. Keys, G.J. Nowacki, C. Carpenter, and W.H. McNab. 2007. Ecological Subregions: Sections and Subsections of the Coterminous United States. USDA Forest Service, General Technical Report WO-76. Washington, DC. 92 pps.

Federal Geographic Data Committee. 2013. Classification of Wetlands and Deepwater Habitats of the United States. FGDC-STD-004-2013. Second Edition. Wetlands Subcommittee, Federal geographic Data Committee and U.S. Fish and Wildlife Service, Washington, D.C. 90 pps.

Flickinger, A. 2010. Iowa Forests Today: An Assessment of the Issues and Strategies for Conserving and Managing Iowa’s Forests. Iowa Department of Natural Resources. 329 pps.

Franzluebbers, A.J., J.A. Stuedemann, H.H. Schomberg, and S.R. Wilkinson. 2000. Soil organic C and N pools under long-term pasture management in the Southern Piedmont USA. Soil Biology and Biochemistry 32:469-478.

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Leake, J., D. Johnson, D. Donnelly, G. Muckle, L. Boddy, and D. Read. 2004. Networks of power and influence: the role of mycorrhizal mycelium in controlling plant communities and agroecosystem functioning. Canadian Journal of Botany 82: 1016-1045.

Millennium Ecosystem Assessment. 2005. Ecosystems and Human Well-Being: Current States and Trends. World Resources Institute. Island Press, Washington, D.C. 948 pages.

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Peel, M.C., B.L. Finlayson, and T.A. McMahon. 2007. Updated world map of the Köppen-Geiger climate classification. Hydrology and Earth System Sciences 11: 1633-1644.

Peterson, C.J. 2000. Catastrophic wind damage to North American forests and the potential impact of climate change. The Science of the Total Environment 262: 287-311.

Pryor, S.C., D. Scavia, C. Downer, M. Gaden, L. Iverson, R. Nordstrom, J. Patz, and G.P. Robertson. 2014. Chapter 18: Midwest. In: J.M. Melillo, T.C. Richmond, and G.W. Yohe, eds. Climate Change Impacts in the United States: The Third National Climate Assessment. U.S. Global Change Research Program, 418-440. Doi:10.7930/J0J1012N.

Schwegman, J.E., G.B. Fell, M. Hutchinson, G. Paulson, W.M. Shepherd, and J. White. 1973. Comprehensive Plan for the Illinois Nature Preserves System, Part 2 The Natural Divisions of Illinois. Illinois Nature Preserves Commission, Rockford, IL. 32 pps.

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Contributors

Lisa Kluesner
Rick Francen

Approval

Suzanne Mayne-Kinney, 12/30/2024

Acknowledgments

This project could not have been completed without the dedication and commitment from a variety of staff members. Team members supported the project by serving on the technical team, assisting with the development of state and community phases of the state-and-transition model, providing peer review and technical editing, and conducting quality control and quality assurance reviews.

List of primary contributors and reviewers.
Organization Name Title Location
Iowa Department of Natural Resources Kevin Andersen State Private Lands Biologist Fairfield, IA
Natural Resources Conservation Service Patrick Chase State Soil Scientist Des Moines, IA
Ron Collman State Soil Scientist Champaign, IL
Tonie Endres Senior Regional Soil Scientist Indianapolis, IN
Rick Francen Soil Scientist Springfield, IL
Lisa Kluesner Ecological Site Specialist Waverly, IA
Jorge, Lugo-Camacho State Soil Scientist Columbia, MO
Kevin Norwood Soil Survey Regional Director Indianapolis, IN
Stanley Sipp Resource Inventory Specialist Champaign, IL
Jason Steele Area Resource Soil Scientist Fairfield, IA
Chris Tecklenberg Acting Regional Ecological Site Specialist Hutchinson, KS
Doug Wallace ACES Ecologist Columbia, MO

Rangeland health reference sheet

Interpreting Indicators of Rangeland Health is a qualitative assessment protocol used to determine ecosystem condition based on benchmark characteristics described in the Reference Sheet. A suite of 17 (or more) indicators are typically considered in an assessment. The ecological site(s) representative of an assessment location must be known prior to applying the protocol and must be verified based on soils and climate. Current plant community cannot be used to identify the ecological site.

Author(s)/participant(s)	Lisa Kluesner
Contact for lead author
Date	01/02/2025
Approved by	Suzanne Mayne-Kinney
Approval date
Composition (Indicators 10 and 12) based on	Annual Production