Natural Resources
Conservation Service
Ecological site F149BY011MA
Well Drained Till Uplands
Last updated: 9/17/2024
Accessed: 11/21/2024
General information
Provisional. A provisional ecological site description has undergone quality control and quality assurance review. It contains a working state and transition model and enough information to identify the ecological site.
MLRA notes
Major Land Resource Area (MLRA): 149B–Long Island-Cape Cod Coastal Lowland
149B—Long Island-Cape Cod Coastal Lowland
This area is in the Embayed Section of the Coastal Plain Province of the Atlantic Plain. It is part of the partially submerged coastal plain of New England. It is mostly an area of nearly level to rolling plains, but it has some steeper hills (glacial moraines). Ridges border the lower plains. The Peconic and Carmans Rivers are on the eastern end of Long Island. The parts of this area in Massachusetts and Rhode Island have no major rivers. This entire area is made up of deep, unconsolidated glacial outwash deposits of sand and gravel. A thin mantle of glacial till covers most of the surface. Some moraines form ridges and higher hills in this area of generally low relief. Sand dunes and tidal marshes are extensive along the coastline.
Classification relationships
USDA-NRCS (USDA, 2006):
Land Resource Region (LRR): S—Northern Atlantic Slope Diversified Farming Region
Major Land Resource Area (MLRA): 149B—Long Island-Cape Cod Coastal Lowland
USDA-FS (Cleland et al., 2007):
Province: 221 Eastern Broadleaf Forest Province
Section: 221A Lower New England
Subsection: 221Ab Cape Cod Coastal Lowland and Islands
Subsection: 221An Long Island Coastal Lowland and Moraine
Ecological site concept
The site consists of very deep, well drained, coarse-loamy soils formed in glacial till. They are on nearly level through moderately steep soils on moraines. Representative soil is Barnstable.
The reference plant community is considered to be an oak-pitch pine forest. Oaks were found to be dominant on terminal moraines within Cape Cod while pitch pine dominated sandy outwash sites (Parshall et al. 2003). Common oaks include black oak, scarlet oak, white, and scrub oak. Pitch pine is subdominant with lesser amounts of beech and hickories. Common shrubs include lowbush blueberry and black huckleberry. Post settlement clearing of forests and an increase in fire has led to a dominance of pitch pine over oaks on terminal moraines within Cape Cod (Parshall et al. 2003). In the absence of severe or frequent fire, oaks replaces pitch pine while other hardwoods such as beech and hickories become a larger component of the forests on more mesic sites.(Parshall et al. 2003).
Associated sites
F149BY010MA |
Moist Till Uplands Moist Till |
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Similar sites
F149BY006NY |
Well Drained Outwash Well-drained outwash |
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Table 1. Dominant plant species
Tree |
(1) Quercus velutina |
---|---|
Shrub |
(1) Vaccinium angustifolium |
Herbaceous |
Not specified |
Physiographic features
The site occurs on nearly level to steep moraines and is not subject to flooding/ponding.
Table 2. Representative physiographic features
Landforms |
(1)
Upland
> Moraine
(2) Ground moraine |
---|---|
Runoff class | Low to very high |
Flooding frequency | None |
Ponding frequency | None |
Elevation | 1,000 ft |
Slope | 35% |
Water table depth | 27 – 72 in |
Aspect | Aspect is not a significant factor |
Climatic features
Coastal regions' climate generally considered maritime, experiences a more moderate climate than inland, i.e., cooler summers and warmer winters and delayed onset of spring. However, coastal regions do experience the brunt of extreme weather such as nor'easters and tropical storms, e.g., hurricanes.
Table 3. Representative climatic features
Frost-free period (characteristic range) | 156-157 days |
---|---|
Freeze-free period (characteristic range) | 199-207 days |
Precipitation total (characteristic range) | 45-49 in |
Frost-free period (actual range) | 156-157 days |
Freeze-free period (actual range) | 197-209 days |
Precipitation total (actual range) | 44-50 in |
Frost-free period (average) | 157 days |
Freeze-free period (average) | 203 days |
Precipitation total (average) | 47 in |
Figure 1. Monthly precipitation range
Figure 2. Monthly minimum temperature range
Figure 3. Monthly maximum temperature range
Figure 4. Monthly average minimum and maximum temperature
Figure 5. Annual precipitation pattern
Figure 6. Annual average temperature pattern
Climate stations used
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(1) HYANNIS [USC00193821], Hyannis, MA
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(2) EAST WAREHAM [USC00192451], East Wareham, MA
Influencing water features
No associated water features
Wetland description
N/A
Soil features
The site consists of moderate to very deep, well drained soils formed in glacial and water deposited materials. Representative soil is Barnstable.
Table 4. Representative soil features
Parent material |
(1)
Till
(2) Glaciofluvial deposits (3) Outwash |
---|---|
Surface texture |
(1) Sandy loam (2) Loamy sand |
Family particle size |
(1) Coarse-loamy over sandy or sandy-skeletal (2) Sandy |
Drainage class | Well drained |
Permeability class | Moderately slow to moderate |
Depth to restrictive layer | 27 – 72 in |
Surface fragment cover <=3" | Not specified |
Surface fragment cover >3" | 9% |
Available water capacity (Depth not specified) |
3 in |
Soil reaction (1:1 water) (Depth not specified) |
3.5 – 6 |
Subsurface fragment volume <=3" (Depth not specified) |
3 – 20% |
Subsurface fragment volume >3" (Depth not specified) |
5 – 10% |
Ecological dynamics
[Caveat: The vegetation information contained in this section and is only provisional, based on concepts, not yet validated with field work.*]
The vegetation groupings described in this section are based on the terrestrial ecological system classification and vegetation associations developed by NatureServe (Comer 2003). Terrestrial ecological systems are specifically defined as a group of plant community types (associations) that tend to co-occur within landscapes with similar ecological processes, substrates, and/or environmental gradients. They are intended to provide a classification unit that is readily mappable, often from terrain and remote imagery, and readily identifiable by conservation and resource managers in the field. A given system will typically manifest itself in a landscape at intermediate geographic scales of tens-to-thousands of hectares and will persist for 50 or more years. A vegetation association is a plant community that is much more specific to a given soil, geology, landform, climate, hydrology, and disturbance history. It is the basic unit for vegetation classification and recognized by the US National Vegetation Classification (US FDGC 2008; USNVC 2017). Each association will be named by the diagnostic and often dominant species that occupy the different height strata (tree, shrub, and herb). Within the NatureServe Explorer database, ecological systems are numbered by a community Ecological System Code (CES) and individual vegetation associations are assigned an identification number called a Community Element Global Code (CEGL).
[*Caveat] The information presented is representative of very complex vegetation communities. Key indicator plants and ecological processes are described to help inform land management decisions. Plant communities will differ across the MLRA because of the naturally occurring variability in weather, soils, and geography. The reference plant community is not necessarily the management goal. The drafts of species lists are merely representative and are not botanical descriptions of all species occurring, or potentially occurring, on this site. They are not intended to cover every situation or the full range of conditions, species, and responses for the site.
This ecological site, set in well-drained till environments, is characterized by a wide mix of plant communities with coastal affinities from Long Island, New York, north to Cape Cod, Massachusetts. These plant communities coincide Northern Atlantic Coastal Plain Dry Oak-Hardwood Forest system(CES203.475), and Northern Atlantic Coastal Plain Heathland and Grassland (CES203.895). Well-drained Till ecological site supports reference plant communities of the oak (Quercus)/heath (Vaccinium spp.) type. oaks (Quercus, spp.). Other canopy hardwoods may include, hickories (Carya spp.), and to a lessor extent, American beech (Fagus gradifolia), and tuliptree (Liriodendron tulipifera) and occasional pines (Pinus spp.). Oaks were found to be dominant on terminal moraines within Cape Cod while pines dominated sandy outwash sites (Parshall et al. 2003). Post settlement clearing of forests and an increase in fire has led to a dominance of pitch pine over oaks on terminal moraines within Cape Cod (Parshall et al. 2003). In the absence of severe or frequent fire, oaks replaces pitch pine while other hardwoods such as beech and hickories become a larger component of the forests on more mesic sites (Parshall et al. 2003). These plant communities coincide with Northern Atlantic Coastal Plain Dry Oak-Hardwood Forest system(CES203.475), and Northern Atlantic Coastal Plain Heathland and Grassland (CES203.895). The prevailing ecological processes are related to coastal influences, such as a coastal climate and storms, and if within close proximity to the coast, maritime effects of wind exposure, salt spray, and sand movement. Fire can be an influence. Threats include development and fragmentation, fire-suppresion, off road vehicles, and invasive plants such as, but not limited to, black locust (Robinia pseudoacacia), tree-of-heaven (Ailanthus altissima), autumn olive (Elaeagnus umbellata), white poplar (Populus alba), multiflora rose (Rosa multiflora), Honeysuckles, (Lonicera spp.), oriental bittersweet (Celatrus orbiculatus), japanese knotweed (Polygonum cuspidatum) wineberry (Rubus phoenicolasius), and Asiatic tearthumb (Polygonum perfoliatum). (Source: NatureServe 2018 [accessed 2019], USNVC 2017 [accessed 2019]).
State and transition model
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State 1 submodel, plant communities
State 2 submodel, plant communities
State 3 submodel, plant communities
State 1
Reference State (Well-drained Till Uplands)
The predominant plant communities of the Well-drained Till ecological site’s Reference State (minimally-managed) include: • Northeastern Coastal Oak / Heath Forest, (Scarlet Oak - Black Oak / Sassafras / Hillside Blueberry Forest), [Quercus coccinea - Quercus velutina / Sassafras albidum / Vaccinium pallidum Forest], - CEGL006375 • Coastal Oak / Mountain Laurel Forest, (Black Oak - Scarlet Oak - Chestnut Oak / Mountain Laurel Forest), [Quercus velutina - Quercus coccinea - Quercus montana / Kalmia latifolia Forest], - CEGL006374 • Or with greater fire frequency: • Pitch Pine - Scarlet Oak Woodland, (Pitch Pine - Scarlet Oak / Hillside Blueberry - (Northern Bayberry) Woodland), [Pinus rigida - Quercus coccinea / Vaccinium pallidum - (Morella pensylvanica) Woodland] - CEGL006381. Other associated coastal/maritime communities can include: • Mid-Atlantic Mesic Mixed Hardwood Forest, (American Beech – Oaks (White Oak, Northern Red Oak) - Tuliptree / (American Holly) Forest, [Fagus grandifolia - Quercus (alba, rubra) - Liriodendron tulipifera / (Ilex opaca) Forest], - CEGL006075 • Northeastern Atlantic Coastal Beech - Oak Forest (American Beech - White Oak - Northern Red Oak Forest), [Fagus grandifolia - Quercus alba - Quercus rubra Forest, - CEGL006377 • Little Bluestem Old-field Meadow, (Little Bluestem - (Broomsedge Bluestem) - Goldenrod species Ruderal Meadow), [Schizachyrium scoparium - (Andropogon virginicus) - Solidago spp. Ruderal Meadow], - CEGL006333 • (Source: NatureServe 2018 [accessed 2019], USNVC 2017 [accessed 2019]).
Community 1.1
Scarlet Oak - Black Oak / Sassafras / Hillside Blueberry Forest
Northeastern Coastal Oak / Heath Forest, (Scarlet Oak - Black Oak / Sassafras / Hillside Blueberry Forest), [Quercus coccinea - Quercus velutina / Sassafras albidum / Vaccinium pallidum Forest], - CEGL006375 This dry coastal forest occurs on rapidly drained, nutrient-poor, sandy or gravelly outwash and till soils. This plant community is dominated by scarlet oak (Quercus coccinea), black oak (Quercus velutina), and occasionally white oak (Quercus alba). Other less abundant canopy associates include chestnut oak (Quercus montana), black birch (Betula lenta), and American holly (Ilex opaca) (usually less than 15% cover). Pitch pine (Pinus rigida) is also common in low cover. Sassafras (Sassafras albidum) can occur in low cover and may indicate influence by coastal (but not maritime) climate. American chestnut (Castanea dentata) saplings may be present. A dense dwarf-shrub heath layer of hillside blueberry (Vaccinium pallidum), lowbush blueberry (Vaccinium angustifolium), and black huckleberry (Gaylussacia baccata) occurs. Blue huckleberry (Gaylussacia frondosa) can sometimes occur. The herbaceous layer is usually sparse, with Pennsylvania sedge (Carex pensylvanica), brakenfern (Pteridium aquilinum), and eastern teaberry (Gaultheria procumbens) being common. Species richness increases with greater canopy gaps, where kinnikinnik (Arctostaphylos uva-ursi), and bushclovers (Lespedeza spp.) occur, and where possibly Canada frostweed (Helianthemum canadense [= Crocanthemum canadense]), Virginia tephrosia [= wild goat’s rue] (Tephrosia virginiana), false foxgloves (Aureolaria spp.), and pinweeds (Lechea spp.) can occur. (Source: NatureServe 2018 [accessed 2019], USNVC 2017 [accessed 2019]). Cross-referenced plant community concepts (typically by political state): BlackOak - Scarlet Oak Woodland (Swain 2016) [MA] Coastal oak-heath forest (Edinger et al. 2014) [NY] Northeastern Coastal Oak-Heath Forest (Sneddon et al. 2010) [Cape Cod National Seashore]
Community 1.2
Black Oak - Scarlet Oak - Chestnut Oak / Mountain Laurel Forest
Coastal Oak / Mountain Laurel Forest, (Black Oak - Scarlet Oak - Chestnut Oak / Mountain Laurel Forest), [Quercus velutina - Quercus coccinea - Quercus montana / Kalmia latifolia Forest], - CEGL006374 This association comprises coastal oak-laurel forests occurring on sandy and gravelly soils. Characteristic dominants are scarlet oak (Quercus coccinea), black oak (Quercus velutina), chestnut oak (Quercus montana), and white oak (Quercus alba). Pines (Pinus) (pitch pine (Pinus rigida), or less comonly white pine (Pinus strobus)) can be present at low cover. Mountain laurel (Kalmia latifolia ) can be the dominant shrub forming extensive, dense patches, with other heath (ericaceous) shrubs such as hillside blueberry (Vaccinium pallidum), lowbush blueberry (Vaccinium angustifolium ), deer blueberry (Vaccinium stamineum), and black huckleberry (Gaylussacia baccata). The herbaceous layer may be sparse due to the dense Kalmia latifolia yet may include braken fern (Pteridium aquilinum), eastern teaberry (Gaultheria procumbens), Pensylvania sedge (Carex pensylvanica), and whitetinge sedge (Carex albicans). With fire suppression, fire-sensitive hardwoods such as American beech (Fagus grandifolia), red maple (Acer rubrum), blackgum (Nyssa sylvatica), and American holly (Ilex opaca )are sometimes present. (Source: NatureServe 2018 [accessed 2019], USNVC 2017 [accessed 2019]). Cross-referenced plant community concepts (typically by political state): BlackOak - Scarlet Oak Woodland (Swain 2016) [MA] Coastal oak-heath forest (Edinger et al. 2014) [NY]
Pathway P1.1A
Community 1.1 to 1.2
Disturbance, greater fire frequency, coastal proximity
Pathway P1.2A
Community 1.2 to 1.1
Succession, reduced fire frequency
State 2
Semi-natural State
Vegetation on lands somewhat conditioned by land use, e.g., managed native plant communities or invasive plant communities.
Community 2.1
Managed Forest Woodland
Community 2.2
Invasive plants
Black locust (Robinia pseudoacacia), Norway Maple (Acer platanoides) or tree-of-heaven (Ailanthus altissima), white poplar (Populus alba), winged burningbush (Euonymus alatus), Japanese knotweed (japanese knotweed (Polygonum cuspidatum), and oriental bittersweet (Celatrus orbiculatus) multiflora rose (Rosa multiflora), bramble wineberry (Rubus phoenicolasius), garlic mustard (Alliaria petiolata), major celindine (Chelidonium majus), ground ivy (Glechoma hederacea), and European lily-of-the-valley (Convallaria majalis), can be present, which may also include native plants. (Source: NatureServe 2018 [accessed 2019], USNVC 2017 [accessed 2019]).
Pathway P2.1A
Community 2.1 to 2.2
Invasive plant establishment
Pathway P2.2A
Community 2.2 to 2.1
Invasive plant management
Conservation practices
Invasive Plant Species Control |
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State 3
Cultural State
Landscapes heavily conditioned by land use, e.g., Plantations/gardens/croplands.
Community 3.1
Plantations / Gardens / Cultivated / Pasture
Transition T1A
State 1 to 2
Disturbance, invasive plant establishment
Conservation practices
Forest Land Management |
---|
Transition T1B
State 1 to 3
Cutting, land clearing, plant establishment, wind erosion control
Conservation practices
Brush Management | |
---|---|
Land Clearing |
Restoration pathway R2A
State 2 to 1
Herbaceous weed treatment, plant removal, plant establishment, successional management
Conservation practices
Brush Management | |
---|---|
Restoration and Management of Natural Ecosystems | |
Native Plant Community Restoration and Management | |
Forest Land Management | |
Invasive Plant Species Control | |
Monitoring and Evaluation |
Transition T2A
State 2 to 3
Cutting, land clearing, plant establishment, wind erosion control
Conservation practices
Land Clearing | |
---|---|
Invasive Plant Species Control | |
Herbaceous Weed Control |
Restoration pathway R3A
State 3 to 1
Herbaceous weed treatment, plant removal, plant establishment, successional management
Conservation practices
Brush Management | |
---|---|
Restoration and Management of Natural Ecosystems | |
Native Plant Community Restoration and Management | |
Invasive Plant Species Control | |
Monitoring and Evaluation | |
Herbaceous Weed Control |
Transition T3A
State 3 to 2
Abandonment, disturbance, invasive plant establishment
Additional community tables
Interpretations
Supporting information
Inventory data references
Site Development and Testing Plan
Future work is needed, as described in a project plan, to validate the information presented in this provisional ecological site description. Future work includes field sampling, data collection and analysis by qualified vegetation ecologists and soil scientists. As warranted, annual reviews of the project plan can be conducted by the Ecological Site Technical Team. A final field review, peer review, quality control, and quality assurance reviews of the ESD are necessary to approve a final document.
References
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Cleland, D.T., J.A. Freeouf, J.E. Keys, G.J. Nowacki, C. Carpenter, and W.H. McNab. 2007. Ecological Subregions: Sections and Subsections of the Coterminous United States. USDA Forest Service, General Technical Report WO-76. Washington, DC. 1–92.
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Comer, P., D. Faber-Langendoen, R. Evans, S. Grawler, C. Josse, G. Kittel, S. Menard, M. Pyne, M. Reid, K. Schultz, K. Snow, and J. Teague. 2003. Ecological Systems of the United States: A Working Classification of U.S. Terrestrial Systems. NatureServe, Arlington, Virginia..
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Edinger, G.J., D.J. Evans, S. Gebauer, T.J. Howard, D. Hunt, and A. Olivero. 2014. Ecological Communities of New York State, Second Edition: A revised and expanded edition of Carol Reschke's Ecological Communities of New York State. New York Natural Heritage Program, New York State Department of Environmental Conservation, Albany, NY..
-
FGDC [Federal Geographic Data Committee]. 2008. National Vegetation Classification Standard, Version 2. Federal Geographic Data Committee, Vegetation Subcommittee, Washington DC..
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Greller, A. 1977. A Classification of Mature Forests on Long Island, New York. Bulletin of the Torrey Botanical Club 104:376–382.
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NatureServe. 2018 (Date accessed). NatureServe Explorer: An online encyclopedia of life [web application]. Version 7.1. NatureServe, Arlington, Virginia. Available http://explorer.natureserve.org.. http://explorer.natureserve.org.
-
Sneddon, L., R. Zaremba, and M. Adams. 2010. Vegetation classification and mapping at Cape Cod National Seashore, Massachusetts.
-
Swain, P.C. 2016. Classification of the natural communities of Massachusetts, Natural Heritage & Endangered Species Program, Massachusetts Division of Fisheries and Wildlife,.
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USNVC [United States National Vegetation Classification]. 2017 (Date accessed). United States National Vegetation Classification Database V2.01. Federal Geographic Data Committee, Vegetation Subcomittee, Washington DC.
Other references
Cleland, D.T., J.A. Freeouf, J.E. Keys, G.J. Nowacki, C. Carpenter, and W.H. McNab. 2007. Ecological Subregions: Sections and Subsections of the Coterminous United States. USDA Forest Service, General Technical Report WO-76. Washington, DC.
Comer, P., D. Faber-Langendoen, R. Evans, S. Gawler, C. Josse, G. Kittel, S. Menard, M. Pyne, M. Reid, K. Schulz, K., Snow, and J.Teague. 2003. Ecological Systems of the United States: A Working Classification of U.S. Terrestrial Systems. NatureServe, Arlington, Virginia.
Edinger, G.J., Evans, D.J., Gebauer, S., Howard, T.G., Hunt, D.M., and A.M. Olivero, A.M. (eds.). 2014. Ecological Communities of New York State, Second Edition: A revised and expanded edition of Carol Reschke's Ecological Communities of New York State. New York Natural Heritage Program, New York State Department of Environmental Conservation, Albany, NY.
FGDC [Federal Geographic Data Committee]. 2008. National Vegetation Classification Standard, Version 2. Federal Geographic Data Committee, Vegetation Subcommittee, Washington DC.
Greller, A. A Classification of Mature Forests on Long Island, New York. Bulletin of the Torrey Botanical Club 104:376–382.
NatureServe 2018. NatureServe Explorer: An online encyclopedia of life [web application]. Version 7.1. NatureServe, Arlington, Virginia. Available http://explorer.natureserve.org. (Accessed: January 2018).
Parshall, T., D.R. Foster, E. Faison, D. MacDonald, and B.C.S. Hansen. 2003. Long‐term history of vegetation and fire in pitch pine–oak forests on cape cod, Massachusetts. Ecology 84:736–748.
Sneddon, L. A., Zaremba, R. E., and M. Adams. 2010. Vegetation classification and mapping at Cape Cod National Seashore, Massachusetts. Natural Resources Technical Report NPS/NER/NRTR--2010/147. National Park Service, Philadelphia, PA.
Swain, P.C. 2016. Classification of the Natural Communities of Massachusetts. Version 2.0. Natural Heritage & Endangered Species Program, Massachusetts Division of Fisheries and Wildlife. Westborough, MA.
United States Department of Agriculture, Natural Resources Conservation Service, 2006. Land Resource Regions and Major land Resource Areas of the United States, the Caribbean, and the Pacific Basin. U.S. Department of Agriculture Handbook 296.
United States Department of Agriculture, Natural Resources Conservation Service, 2015. National Soils Information System (NASIS).
USNVC [United States National Vegetation Classification]. 2017. United States National Vegetation Classification Database, V2.01. Federal Geographic Data Committee, Vegetation Subcommittee, Washington DC. http://usnvc.org/explore-classification/ (Accessed: 2018)
Contributors
Nels Barrett, Ph.D.
Joshua Hibit
Approval
Nels Barrett, 9/17/2024
Rangeland health reference sheet
Interpreting Indicators of Rangeland Health is a qualitative assessment protocol used to determine ecosystem condition based on benchmark characteristics described in the Reference Sheet. A suite of 17 (or more) indicators are typically considered in an assessment. The ecological site(s) representative of an assessment location must be known prior to applying the protocol and must be verified based on soils and climate. Current plant community cannot be used to identify the ecological site.
Author(s)/participant(s) | |
---|---|
Contact for lead author | |
Date | 05/23/2020 |
Approved by | Nels Barrett |
Approval date | |
Composition (Indicators 10 and 12) based on | Annual Production |
Indicators
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Number and extent of rills:
-
Presence of water flow patterns:
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Number and height of erosional pedestals or terracettes:
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Bare ground from Ecological Site Description or other studies (rock, litter, lichen, moss, plant canopy are not bare ground):
-
Number of gullies and erosion associated with gullies:
-
Extent of wind scoured, blowouts and/or depositional areas:
-
Amount of litter movement (describe size and distance expected to travel):
-
Soil surface (top few mm) resistance to erosion (stability values are averages - most sites will show a range of values):
-
Soil surface structure and SOM content (include type of structure and A-horizon color and thickness):
-
Effect of community phase composition (relative proportion of different functional groups) and spatial distribution on infiltration and runoff:
-
Presence and thickness of compaction layer (usually none; describe soil profile features which may be mistaken for compaction on this site):
-
Functional/Structural Groups (list in order of descending dominance by above-ground annual-production or live foliar cover using symbols: >>, >, = to indicate much greater than, greater than, and equal to):
Dominant:
Sub-dominant:
Other:
Additional:
-
Amount of plant mortality and decadence (include which functional groups are expected to show mortality or decadence):
-
Average percent litter cover (%) and depth ( in):
-
Expected annual annual-production (this is TOTAL above-ground annual-production, not just forage annual-production):
-
Potential invasive (including noxious) species (native and non-native). List species which BOTH characterize degraded states and have the potential to become a dominant or co-dominant species on the ecological site if their future establishment and growth is not actively controlled by management interventions. Species that become dominant for only one to several years (e.g., short-term response to drought or wildfire) are not invasive plants. Note that unlike other indicators, we are describing what is NOT expected in the reference state for the ecological site:
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Perennial plant reproductive capability:
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