Ecological dynamics
[Caveat: The vegetation information contained in this section and is only provisional, based on concepts, not yet validated with field work.*]
The vegetation groupings described in this section are based on the terrestrial ecological system classification and vegetation associations developed by NatureServe (Comer 2003). Terrestrial ecological systems are specifically defined as a group of plant community types (associations) that tend to co-occur within landscapes with similar ecological processes, substrates, and/or environmental gradients. They are intended to provide a classification unit that is readily mappable, often from terrain and remote imagery, and readily identifiable by conservation and resource managers in the field. A given system will typically manifest itself in a landscape at intermediate geographic scales of tens-to-thousands of hectares and will persist for 50 or more years. A vegetation association is a plant community that is much more specific to a given soil, geology, landform, climate, hydrology, and disturbance history. It is the basic unit for vegetation classification and recognized by the US National Vegetation Classification (US FDGC 2008; USNVC 2017). Each association will be named by the diagnostic and often dominant species that occupy the different height strata (tree, shrub, and herb). Within the NatureServe Explorer database, ecological systems are numbered by a community Ecological System Code (CES) and individual vegetation associations are assigned an identification number called a Community Element Global Code (CEGL).
[*Caveat] The information presented is representative of very complex vegetation communities. Key indicator plants and ecological processes are described to help inform land management decisions. Plant communities will differ across the MLRA because of the naturally occurring variability in weather, soils, and geography. The reference plant community is not necessarily the management goal. The drafts of species lists are merely representative and are not botanical descriptions of all species occurring, or potentially occurring, on this site. They are not intended to cover every situation or the full range of conditions, species, and responses for the site.
The Well-drained Outwash ecological site is characterized by a wide mix of xeric plant communities with coastal affinities from Long Island, New York, north to Cape Cod, Massachusetts. These plant communities coincide with Northern Atlantic Coastal Pitch Pine Barrens system (CES203.269), Northern Atlantic Coastal Plain Dry Oak-Hardwood Forest system (CES203.475), and Northern Atlantic Coastal Plain Heathland and Grassland (CES203.895). The prevailing ecological processes are related to coastal influences, such as a coastal climate and storms, and if within close proximity to the coast, maritime effects of wind exposure, salt spray, and sand movement. This ecological site is influenced by fire. A decadal fire frequency maintains the character of pine barrens, dominated by pitch Pine (Pinus rigida), as well as some heathlands and grasslands with characteristic shrubs Northern bayberry (Morella pensylvanica) kinnikinnik (Arctostaphylos uva-ursi), and goldenheathers (Hudsonia spp.) with grasses little bluestem (Schizachyrium scoparium) or coastal little bluestem (Schizachyrium littorale). Longer fire intervals and/or cold-air drainage (frostpockets) will support various oaks (Quercus), predominately scrub oak (Q. ilicifolia), dwarf chesnut oak (Q. prinoides), scarlet oak (Q. coccinea), black oak (Q. velutina), as well as white oak (Q. alba) and chesnut oak (Q. montana). Threats include development and fragmentation, fire-suppresion, off road vehicles, and invasive plants such as, but not limited to, Morrow’s honeysuckle, (Lonicera morowii), oriental bittersweet (Celatrus orbiculatus), wineberry (Rubus phoenicolasius) tree-of-heaven (Ailanthus altissima). (Source: NatureServe 2018 [accessed 2019], USNVC 2017 [accessed 2019]).
State 1
Reference State (Well-drained Outwash)
The predominant plant communities of the Well-drained Outwash ecological site Reference State (minimally-managed) include:
• Northeastern Coastal Oak / Heath Forest, (Scarlet Oak - Black Oak / Sassafras / Hillside Blueberry Forest), [Quercus coccinea - Quercus velutina / Sassafras albidum / Vaccinium pallidum Forest], - CEGL006375
Or with greater fire frequency:
• Pitch Pine - Scarlet Oak Woodland, (Pitch Pine - Scarlet Oak / Hillside Blueberry - (Northern Bayberry) Woodland), [Pinus rigida - Quercus coccinea / Vaccinium pallidum - (Morella pensylvanica) Woodland] - CEGL006381.
Other associated coastal/maritime communities can include:
• Coastal Oak / Mountain Laurel Forest, (Black Oak - Scarlet Oak - Chestnut Oak / Mountain Laurel Forest), [Quercus velutina - Quercus coccinea - Quercus montana / Kalmia latifolia Forest], - CEGL006374
• Northeastern Atlantic Coastal Beech - Oak Forest, (American Beech - White Oak - Northern Red Oak Forest), [Fagus grandifolia - Quercus alba - Quercus rubra Forest], - CEGL006377
• Northern Tall Maritime Scrub Forest, (Canadian Serviceberry - Viburnum species - Northern Bayberry Scrub Forest), [Amelanchier canadensis - Viburnum spp. - Morella pensylvanica Scrub Forest], - CEGL006379
• Northeastern Maritime Forest, (Black Cherry - Sassafras - Canadian Serviceberry - Black Oak / Roundleaf Greenbrier Forest0, [Prunus serotina - Sassafras albidum - Amelanchier canadensis - Quercus velutina / Smilax rotundifolia Forest], - CEGL006145
• Maritime Red-cedar Woodland, (Eastern Red-cedar / Northern Bayberry Woodland), [Juniperus virginiana / Morella pensylvanica Woodland], - CEGL006212
• Long Island Maritime Beech Forest, (American Beech / Roundleaf Greenbrier Forest), [Fagus grandifolia / Smilax rotundifolia Forest], - CEGL006043
• Sandplain Heathland, (Black Huckleberry - Lowbush Blueberry - Bearberry / Shore Little Bluestem Dwarf-shrubland), [Gaylussacia baccata - Vaccinium angustifolium - Arctostaphylos uva-ursi / Schizachyrium littorale Dwarf-shrubland], - CEGL006066
• Sandplain Grassland, (Northern Bayberry / Shore Little Bluestem - Poverty Oatgrass Shrub Grassland), [Morella pensylvanica / Schizachyrium littorale - Danthonia spicata Shrub Grassland], - CEGL006067
• Hempstead Plain Grassland, (Little Bluestem - Indiangrass - Common Goldstar - Horseflyweed Grassland), [Schizachyrium scoparium - Sorghastrum nutans - Hypoxis hirsuta - Baptisia tinctoria Grassland], - CEGL006187
• Little Bluestem Old-field Meadow, (Little Bluestem - (Broomsedge Bluestem) - Goldenrod species Ruderal Meadow), [Schizachyrium scoparium - (Andropogon virginicus) - Solidago spp. Ruderal Meadow], - CEGL006333
(Source: NatureServe 2018 [accessed 2019], USNVC 2017 [accessed 2019])
Community 1.1
Scarlet Oak - Black Oak / Sassafras / Hillside Blueberry Forest
Northeastern Coastal Oak / Heath Forest, (Scarlet Oak - Black Oak / Sassafras / Hillside Blueberry Forest), [Quercus coccinea - Quercus velutina / Sassafras albidum / Vaccinium pallidum Forest], - CEGL006375
This dry coastal forest occurs on rapidly drained, nutrient-poor, sandy or gravelly outwash and till soils. This plant community is dominated by scarlet oak (Quercus coccinea), black oak (Quercus velutina), and occasionally white oak (Quercus alba). Other less abundant canopy associates include chestnut oak (Quercus montana), black birch (Betula lenta), and American holly (Ilex opaca) (usually less than 15% cover). Pitch pine (Pinus rigida) is also common in low cover. Sassafras (Sassafras albidum) can occur in low cover and may indicate influence by coastal (but not maritime) climate. American chestnut (Castanea dentata) saplings may be present. A dense dwarf-shrub heath layer of hillside blueberry (Vaccinium pallidum), lowbush blueberry (Vaccinium angustifolium), and black huckleberry (Gaylussacia baccata) occurs. Blue huckleberry (Gaylussacia frondosa) can sometimes occur. The herbaceous layer is usually sparse, with Pennsylvania sedge (Carex pensylvanica), brakenfern (Pteridium aquilinum), and eastern teaberry (Gaultheria procumbens) being common. Species richness increases with greater canopy gaps, where kinnikinnik (Arctostaphylos uva-ursi), and bushclovers (Lespedeza spp.) occur, and where possibly Canada frostweed (Helianthemum canadense [= Crocanthemum canadense]), Virginia tephrosia [= wild goat’s rue] (Tephrosia virginiana), false foxgloves (Aureolaria spp.), and pinweeds (Lechea spp.) can occur. (Source: NatureServe 2018 [accessed 2019], USNVC 2017 [accessed 2019]).
Cross-referenced plant community concepts (typically by political state):
BlackOak - Scarlet Oak Woodland (Swain 2016) [MA]
Coastal oak-heath forest (Edinger et al. 2014) [NY]
Northeastern Coastal Oak-Heath Forest (Sneddon et al. 2010) [Cape Cod National Seashore]
Community 1.2
Pitch Pine - Scarlet Oak / Hillside Blueberry - (Northern Bayberry) Woodland
Pitch Pine - Scarlet Oak Woodland, (Pitch Pine - Scarlet Oak / Hillside Blueberry - (Northern Bayberry) Woodland), [Pinus rigida - Quercus coccinea / Vaccinium pallidum - (Morella pensylvanica) Woodland] - CEGL006381.
This vegetation is a matrix woodland/low forest comprising the pine-barren lands from Long Island, New York, north to Cape Cod, Massachusetts. Typically, it occurs on very well-drained sandy outwash. The canopy is largely pitch pine (Pinus rigida) plus an admixture of oaks (Quercus): scarlet oak (Q. coccinea), black oak (Q. velutina), dwarf chestnut oak (Q. prinoides), white oak (Q. alba), and northern red oak (Q. rubra) and occasionally post oak (Q. stellata). Tall shrubs are dominated by bear or scrub oak (Quercus ilicifolia) can be sporadic to locally well-developed and northern bayberry (Morella pensylvanica) can also commonly occur. Heaths tend to form a dense dwarf-shrub layer, especially black huckleberry (Gaylussacia baccata), lowbush blueerry (Vaccinium angustifolium), and hillside blueberry (Vaccinium pallidum). The herb layer is often sparse and characterized by brackenfern (Pteridium aquilinum), eastern teaberry (Gaultheria procumbens), and wavy hairgrass (Deschampsia flexuosa). (Source: NatureServe 2018 [accessed 2019], USNVC 2017 [accessed 2019]).
Cross-referenced plant community concepts (typically by political state):
Coastal Forest/Woodland (Swain 2016) [MA]
Pitch pine – oak Forest (Edinger et al. 2014) [NY]
Pitch pine – oak Forest (Sneddon et al. 2010) [Cape Cod National Seashore]
Pathway CP1.1-1.2
Community 1.1 to 1.2
disturbance, greater fire frequency, coastal proximity
Pathway CP1.2-1.1
Community 1.2 to 1.1
succession, lessor fire frequency
State 2
Semi-natural State
Vegetation on lands somewhat conditioned by land use, e.g., managed native plant communities or invasive plant communities.
Community 2.1
Managed Forest Woodland
Community 2.2
Black Locust Ruderal Forest
Ruderal Black Locust Forest, (Black Locust Ruderal Forest), [Robinia pseudoacacia Ruderal Forest], - CEGL007279
Black locust (Robinia pseudoacacia) can establish on abandoned croplands, pasturing, and former homesites. Associated plants can vary from site to site and include black cherry (Prunus serotina), red ceadr (Juniperus virginiana), American elm (Ulmus americana), slippery elm (Ulmus rubra), shagbark hickory (Carya ovata), hackberry (Celtis occidentalis), black walnut (Juglans nigra), red oak (Quercus rubra), and in some areas Norway Maple (Acer platanoides) or tree-of-heaven (Ailanthus altissima). The understory vegetation is also highly variable depending on site history and often includes (poisen ivy) Toxicodendron radicans; northern spicebush (Lindera benzoin) is sometimes present in loamy , more mesic conditions. The invasive non-native shrub multiflora rose (Rosa multiflora) and bramble wineberry (Rubus phoenicolasius) can be present. Other non-native plants like, garlic mustard (Alliaria petiolata), major celindine (Chelidonium majus), ground ivy (Glechoma hederacea), and European lily-of-the-valley (Convallaria majalis), can characterize the herb layer, which may also include native plants. (Source: NatureServe 2018 [accessed 2019], USNVC 2017 [accessed 2019]).
Other non-native plants can include white poplar (Populus alba), winged burningbush (Euonymus alatus), Japanese knotweed (japanese knotweed (Polygonum cuspidatum), and oriental bittersweet (Celatrus orbiculatus)
Cross-referenced plant community concepts (typically by political state):
Successional southern hardwoods (Edinger et al. 2014) [NY]
Black Locust successional forest (Sneddon et al. 2010) [Cape Cod National Seashore]
Pathway CP2.1-2.1
Community 2.1 to 2.2
Invasive Plant establishment
Pathway CP2.2-2.1
Community 2.2 to 2.1
Invasive Plant Management
Invasive Plant Species Control |
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State 3
Cultural State
Landscapes heavily conditioned by land use, e.g., Plantations/gardens/croplands.
Community 3.1
Orchard Grass - Timothy - Fescue species - Goldenrod species Herbaceous Vegetation
Northeastern Old Field Measdow, (Orchard Grass - Timothy - Fescue species - Goldenrod species Herbaceous Vegetation), [Dactylis glomerata - Phleum pratense - Festuca spp. - Solidago spp. Herbaceous Vegetation], - CEGL006107
This is a broadly defined vegetation type of pastures and is largely composed of non-native cool-season grasses and herbs (generally of European origin) in the early stages of succession. The fields are typically mowed at least annually. Species composition varies from site to site, from site to site but generally feature Orchard Grass (Dactylis glomerata). Timothy (Phleum pratense)- Fescue grasses (Festuca spp.), and some goldenrods (Solidago spp.). Other graminoid associates may include creeping bentgrass (Agrostis stolonifera), winer bentgrass (Agrostis hyemalis), creeping wildrye (Elymus repens), smooth brome (Bromus inermis), cheatgrass (Bromus tectorum), perennial ryegrass (Lolium perenne), Kentucky bluegrass (Poa pratensis), Canada bluegrass (Poa compressa), and sweet vernalgrass (Anthoxanthum odoratum). Forbs scattered among the grasses are varied but include hawkweeds (Hieracium spp.), yellow oxalis (Oxalis stricta), common yarrow (Achillea millefolium), common milkweed (Asclepias syriaca), wrinkleleaf goldenrod (Solidago rugosa), gray goldenrod (Solidago nemoralis), early goldenrod (Solidago juncea), Canada goldenrod (Solidago canadensis), tall goldenrod (Solidago altissima), flattop goldenrod (Euthamia graminifolia), field chickweed (Cerastium arvense), evening primrose (Oenothera biennis), common cinquefoil (Potentilla simplex), calico American-aster (Symphyotrichum lateriflorum), New England American-aster (Symphyotrichum novae-angliae), white panicle American -aster (Symphyotrichum lanceolatum), Queen Anne’s lace (Daucus carota), annual agweed (Ambrosia artemisiifolia), bird vetch (Vicia cracca), sweetclover (Trifolium spp.), and many others (Source: NatureServe 2018 [accessed 2019], USNVC 2017 [accessed 2019]).
Cross-referenced plant community concepts (typically by political state):
Cultural Grassland (Swain Swain 2016) [MA]
Successional Old Field (Edinger et al. 2014) [NY]
Nothestern Old Field (Sneddon et al. 2010) [Cape Cod National Seashore]
Transition T1-2
State 1 to 2
disturbance, invasive plant establishment
Transition T1-3
State 1 to 3
cutting, land clearing, plant establishment
Brush Management |
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Land Clearing |
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Restoration pathway R2-1
State 2 to 1
herbaceous weed treatment, plant removal, plant establishment, successional management
Brush Management |
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Restoration and Management of Natural Ecosystems |
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Native Plant Community Restoration and Management |
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Forest Land Management |
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Invasive Plant Species Control |
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Monitoring and Evaluation |
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Transition T2-3
State 2 to 3
cutting, land clearing, plant establishment
Land Clearing |
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Invasive Plant Species Control |
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Herbaceous Weed Control |
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Restoration pathway R3-1
State 3 to 1
herbaceous weed treatment, plant removal, plant establishment, successional management
Brush Management |
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Restoration and Management of Natural Ecosystems |
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Native Plant Community Restoration and Management |
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Invasive Plant Species Control |
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Monitoring and Evaluation |
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Herbaceous Weed Control |
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Transition T3-2
State 3 to 2
disturbance, invasive plant establishment