Ecological dynamics
Major Successional Stages of Forestland Development
HERBACEOUS: Vegetation is dominated by grasses and forbs under full sunlight. This stage is experienced after a major disturbance such as wildfire. Skeleton forest (dead trees) remaining after fire or other disturbances have little or no affect on the composition and production of the herbaceous vegetation.
SHRUB-HERBACEOUS: Herbaceous vegetation and woody shrubs dominate the site. This stage is experienced within two to three years after fire or other major disturbance. Various amounts of tree seedlings (less than 20 inches (51 cms) in height) may be present up to the point where they are obviously a major component of the vegetal structure.
SAPLING: In the absence of disturbance, the tree seedlings develop into saplings (20 inches to 4.5 feet (51 cms to 137 cms) in height) with a canopy cover generally of about 5 to 10 percent. Vegetation consists of grasses, forbs and shrubs in association with tree saplings.
IMMATURE FORESTLAND: The visual aspect and vegetal structure are dominated by ponderosa pine greater than 4.5 feet in height. The upper crown of dominant and co-dominant ponderosa pine are cone or pyramidal shaped. Seedlings and saplings of pines are common in the understory. Dominants are the tallest trees on the site; co-dominants are 65 to 85 percent of the highest of dominant trees. Understory vegetation is moderately influenced by a tree overstory canopy of about 5 to 15 percent.
MATURE FORESTLAND: The visual aspect and vegetal structure are dominated by ponderosa pine that have reached or are near maximal heights for the site. Tree canopy cover ranges from 10 to 20 percent. This stage of community development is assumed to be representative of this forestland site in the pristine environment.
OVER-MATURE FORESTLAND: In the absence of wildfire or other major disturbances the tree canopy will continue to expand. This stage is dominated by ponderosa pine that have reached maximal heights for the site. Understory vegetation is sparse or absent due to tree competition, overstory shading, duff accumulation, etc. Few seedlings or saplings of ponderosa pine are found in the understory. Tree canopy cover is commonly greater than 20 percent.
Ponderosa pine is shade intolerant and grows most rapidly in near full sunlight. A strong tendency exists for climax pine stands to be even- aged in small groups rather than being truly uneven-aged.
The successional status of ponderosa pine can be expressed in terms of its successional role, which ranges from seral to climax depending on specific site conditions. It plays a climax role on sites toward the extreme limits of its environmental range and becomes increasingly seral with more favorable conditions. On sites with more favorable moisture, pine encounters greater competition and must establish itself opportunistically. On moist sites it is usually seral. On severe sites it is climax by default because other species cannot establish. On such sites, establishment is likely to be highly dependent upon the cyclical nature of large seed crops and favorable weather conditions.
Fire Ecology:
Historically, surface fires were most common in Ponderosa pine stands, with occasional flare-ups occurring in brush patches. Crown fires were unlikely in most stands. Long, loosely packed pine needles and herbaceous species maintained frequent and mild surface burns. These fires created openings for pine seedling establishment, thus maintaining its persistence. These fires also thinned saplings and maintained the relatively open understories. The recent increase in understory density has been attributed to the current longer fire intervals. Ponderosa pine is fire-adapted in all stages of its life history and is especially well adapted to light, regular surface fires. The seeds prefer openings with mineral soil usually prepared by fire for seedbed. For the seedlings and young trees, early development of insulative bark, shielded meristems, high moisture content in living needles, and rapid extension of taproots reduce their mortality from fire. For mature trees, thick bark, deep roots, and low- flammability crown structures help them survive most fires. Following a stand-replacement fire, the successional sequence in these types’ proceeds from herbaceous species to shrub and hardwood, and finally ponderosa pine stages. Sedge is top-killed by fire, with rhizomes protected by insulating soil. The rhizomes of sedge species may be killed by high-severity fires that remove most of the soil organic layer. Reestablishment after fire occurs by seed establishment, rhizomatous spread or both. Desert needlegrass has persistent dead leaf bases, which make it susceptible to burning. Fire removes the accumulation; a rapid, cool fire will not burn deep into the root crown.
Bottlebrush squirreltail's small size, coarse stems, and sparse leafy material aid in its tolerance of fire. Postfire regeneration occurs from surviving root crowns and from on- and off-site seed sources. Frequency of disturbance greatly influences postfire response of bottlebrush squirreltail. Undisturbed plants within a 6 to 9 year age class generally contain large amounts of dead material, increasing bottlebrush squirreltail's susceptibility to fire.
State 1
Reference State
The Reference State 1.0 is a representative of the natural range of variability under pristine conditions. This site has four general community phases: (1) a mature woodland phase, (2) a sucker/sapling phase, (3) an immature woodland phase and (4) an over mature woodland/conifer phase. State dynamics are maintained by interactions between climatic patterns and disturbance regimes. Negative feedbacks enhance ecosystem resilience and contribute to the stability of the state. These include the presence of all structural and functional groups, fuel loads, and retention of organic matter and nutrients. Plant community phase changes are primarily driven by fire, periodic drought, insect or disease attack.
Community 1.1
Mature Forestland
The reference plant community is dominated by ponderosa pine. Altered andesite buckwheat is the principal understory plant. An overstory canopy cover of about 15 percent is assumed to be representative of tree dominance on this site in the pristine environment. Overstory tree canopy composition is typically 100 percent ponderosa pine. Singleleaf piñyon, Utah juniper and/or Jeffrey pine may compose up to 5 percent of the overstory.
MATURE FORESTLAND: The visual aspect and vegetal structure are dominated by ponderosa pine that have reached or are near maximal heights for the site. Tree canopy cover ranges from 10 to 20 percent. This stage of community development is assumed to be representative of this forestland site in the pristine environment.
Forest overstory. MATURE FORESTLAND: The visual aspect and vegetal structure are dominated by ponderosa pine that have reached or are near maximal heights for the site. Tree canopy cover ranges from 10 to 20 percent. This stage of community development is assumed to be representative of this forestland site in the pristine environment.
Forest understory. Understory vegetative composition is about 30 percent grasses, 05 percent forbs and 65 percent shrubs and young trees when the average overstory canopy is medium (10 to 20 percent). Average understory production ranges from 25 to 125 pounds per acre with a medium canopy cover. Understory production includes the total annual production of all species within 4.5 feet of the ground surface.
Table 5. Annual production by plant type
| Plant type |
Low (lb/acre) |
Representative value (lb/acre) |
High (lb/acre) |
| Tree |
9 |
18 |
44 |
| Shrub/Vine |
8 |
15 |
38 |
| Grass/Grasslike |
7 |
14 |
37 |
| Forb |
1 |
3 |
6 |
| Total |
25 |
50 |
125 |
Community 1.2
Herbaceous/Shrub
HERBACEOUS: Vegetation is dominated by grasses and forbs under full sunlight. This stage is experienced after a major disturbance such as wildfire. Skeleton forest (dead trees) remaining after fire or other disturbances have little or no affect on the composition and production of the herbaceous vegetation.
SHRUB-HERBACEOUS: Herbaceous vegetation and woody shrubs dominate the site. This stage is experienced within two to three years after fire or other major disturbance. Various amounts of tree seedlings (less than 20 inches (51 cms) in height) may be present up to the point where they are obviously a major component of the vegetal structure.
Community 1.3
Sapling to Immature Forest
SAPLING: In the absence of disturbance, the tree seedlings develop into saplings (20 inchesto 4.5 feet (51 cms to 137 cms) in height) with a canopy cover generally of about 5 to 10 percent. Vegetation consists of grasses, forbs and shrubs in association with tree saplings.
IMMATURE FORESTLAND: The visual aspect and vegetal structure are dominated by ponderosa pine greater than 4.5 feet in height. The upper crown of dominant and co-dominant ponderosa pine are cone or pyramidal shaped. Seedlings and saplings of pines are common in the understory. Dominants are the tallest trees on the site; co-dominants are 65 to 85 percent of the highest of dominant trees. Understory vegetation is moderately influenced by a tree overstory canopy of about 5 to 15 percent.
Community 1.4
Over-Mature Forest
OVER-MATURE FORESTLAND: In the absence of wildfire or other major disturbances the tree canopy will continue to expand. This stage is dominated by ponderosa pine that have reached maximal heights for the site. Understory vegetation is sparse or absent due to tree
competition, overstory shading, duff accumulation, etc. Few seedlings or saplings of ponderosa pine are found in the understory. Tree canopy cover is commonly greater than 20 percent.
Pathway 1.1a
Community 1.1 to 1.2
This pathway is a result of fire which reduces trees and increases herbaceous vegetation.
Pathway 1.1b
Community 1.1 to 1.4
This pathway is a result of time and lack of disturbance.
Pathway 1.2a
Community 1.2 to 1.3
This pathway is a result of time and lack of disturbance.
Pathway 1.3a
Community 1.3 to 1.1
This pathway is a result of time and lack of disturbance.
Pathway 1.3b
Community 1.3 to 1.2
This pathway is a result of disturbance such as fire, insects, or disease.
Pathway 1.4a
Community 1.4 to 1.2
This pathway is a result of disturbance, such as fire.
State 2
Current Potential State
This state is similar to the Reference State 1.0 with four similar community phases. Ecological function has not changed, however the resiliency of the state has been reduced by the presence of invasive weeds. Non-natives may increase in abundance but will not become dominant within this State. These non-natives can be highly flammable and can promote fire where historically fire had been infrequent. Negative feedbacks enhance ecosystem resilience and contribute to the stability of the state. These feedbacks include the presence of all structural and functional groups, low fine fuel loads, and retention of organic matter and nutrients. Positive feedbacks decrease ecosystem resilience and stability of the state. These include the non-natives’ high seed output, persistent seed bank, rapid growth rate, ability to cross pollinate, and adaptations for seed dispersal.
Community 2.1
Mature forest with non-native plants
This community is similar to 1.1 with non-native plants.
The visual aspect and vegetal structure are dominated by ponderosa pine that have reached or are near maximal heights for the site. Tree canopy cover ranges from 10 to 20 percent. This stage of community development is assumed to be representative of this forestland site in the pristine environment.
Community 2.2
Herbaceous/Shrub/non-native plants
This community is similar to 1.2 with non-native plants.
Vegetation is dominated by grasses and forbs under full sunlight. This stage is experienced after a major disturbance such as wildfire. Skeleton forest (dead trees) remaining after fire or other disturbances have little or no affect on the composition and production of the herbaceous vegetation.
SHRUB-HERBACEOUS: Herbaceous vegetation and woody shrubs dominate the site. This stage is experienced within two to three years after fire or other major disturbance. Various amounts of tree seedlings (less than 20 inches (51 cms) in height) may be present up to the point where they are obviously a major component of the vegetal structure.
Community 2.3
Sapling to Immature Forest
This community is similar to 1.3 with non-native plants.
SAPLING: In the absence of disturbance, the tree seedlings develop into saplings (20 inches to 4.5 feet (51 cms to 137 cms) in height) with a canopy cover generally of about 5 to 10 percent. Vegetation consists of grasses, forbs and shrubs in association with tree saplings.
IMMATURE FORESTLAND: The visual aspect and vegetal structure are dominated by ponderosa pine greater than 4.5 feet in height. The upper crown of dominant and co-dominant ponderosa pine are cone or pyramidal shaped. Seedlings and saplings of pines are common in the understory. Dominants are the tallest trees on the site; co-dominants are 65 to 85 percent of the highest of dominant trees. Understory vegetation is moderately influenced by a tree overstory canopy of about 5 to 15 percent.
Community 2.4
Over-Mature Forest with non-native plants
This community is similar to 1.4 with non-native plants. In the absence of wildfire or other major disturbances the tree canopy will continue to expand. This stage is dominated by ponderosa pine that have reached maximal heights for the site. Understory vegetation is sparse or absent due to tree competition, overstory shading, duff accumulation, etc. Few seedlings or saplings of ponderosa pine are found in the understory. Tree canopy cover is commonly greater than 20 percent.
Pathway 2.1a
Community 2.1 to 2.2
This pathway is a result of disturbance (fire) that removes tree canopy and increases understory grasses and shrubs.
Pathway 2.1b
Community 2.1 to 2.4
This pathway is a result of time and lack of disturbance.
Pathway 2.2a
Community 2.2 to 2.3
This pathway is a result of time and lack of disturbance.
Pathway 2.3a
Community 2.3 to 2.1
This pathway is a result of time and lack of disturbance.
Pathway 2.3b
Community 2.3 to 2.2
This pathway is a result of disturbance, such as fire, that removes trees and increases grasses and shrubs.
Pathway 2.4a
Community 2.4 to 2.2
This pathway is a result of disturbance, such as fire, that removes tree canopy.
Transition T1A
State 1 to 2
This transition occurs with establishment of non-native plants.