Physiographic features

This site typically occurs on sedimentary plains uplands where it occurs on swales, rises, ridges, and hills. The site also occurs on areas of outwash and on remnant till which is mantled with moderately coarse-textured, eolian deposits. Slope ranges from 0 to 50 percent.

Table 2. Representative physiographic features

Climatic features

MLRA 54 is considered to have a continental climate with cold winters and hot summers, low humidity, light rainfall, and much sunshine. Extremes in temperature are common and characteristic of MLRA 54. The continental climate is the result of this MLRA’s location in the geographic center of North America. There are few natural barriers on the northern Great Plains, so air masses move unobstructed across the plains and account for rapid changes in temperature.

Annual precipitation ranges from 14 to 18 inches per year. The normal average annual temperature is about 42° F. January is the coldest month with average temperatures ranging from about 13° F (Beach, ND) to about 16° F (Bison, SD). July is the warmest month with temperatures averaging from about 69° F (Beach, ND) to about 72° F (Timber Lake, SD). The range of normal average monthly temperatures between the coldest and warmest months is about 57° F. This large temperature range attests to the continental nature of MLRA 54’s climate. Wind speeds average about 11 miles per hour, ranging from about 13 miles per hour during the spring to about 10 miles per hour during the summer. Daytime wind speeds are generally stronger than nighttime wind speeds, and occasional strong storms may bring brief periods of high winds with gusts to more than 50 miles per hour.

Growth of native cool-season plants begins in late March and continues through early to mid-July. Native warm-season plants begin growth in mid-May and continue to the end of August. Green-up of cool-season plants can occur again in September and October when adequate soil moisture is present.

Table 3. Representative climatic features

Climate stations used

• (1) FT YATES 4 SW [USC00323207], Fort Yates, ND

• (2) DUPREE [USC00392429], Dupree, SD

• (3) HETTINGER [USC00324178], Hettinger, ND

• (4) WATFORD CITY [USC00329233], Watford City, ND

• (5) MANDAN EXP STN [USC00325479], Mandan, ND

• (6) LUDLOW 3 SSE [USC00395048], Ludlow, SD

• (7) HEBRON [USC00324102], Hebron, ND

Influencing water features

This site does not receive significant additional water, either as runoff from adjacent slopes or from a seasonal high-water table. Depth to the water table typically exceeds 6 feet throughout the growing season, but it may be as shallow as 5 feet in early spring of some years. Surface infiltration is typically moderately rapid. Permeability is moderately rapid in the upper part of the soil. Where a contrasting texture is present below a depth of 20 inches, permeability ranges from moderately slow to moderate. Water loss is through percolation below the root zone and evapotranspiration.

Salinization Risk – Removal of perennial vegetation (e.g., annual cropping), above this ecological site (recharge area), results in increased water moving down through the soil profile. The potential salinization of the site can occur in any of the vegetative states; however, it is most prevalent in the State 4.0 Go-Back. This downward water movement leaches salts over time creating shallow saline groundwater immediately above a less permeable layer. Due to gravity, water moves downward through the soil profile then laterally through a porous layer, such as a coal or sand/gravel seam, transporting salts to the discharge area. Below the porous layer, is a less permeable layer such as soft sedimentary bedrock. Where shallow saline groundwater occurs, salts often concentrate at or near the soil surface through capillary rise (discharge area). In capillary rise, water moves from where the soil is saturated, or nearly so, to drier soil against the force of gravity. Evaporation at the soil surface dries the soil and “pulls” water by capillary flow from the wet soil zone. Because only pure water evaporates, salts are left behind.

Figure 8. Increased salinity levels will make establishment of grasses and forbs more difficult. Treatment of the recharge area is a requirement to reclaim these soils from salinization. Some salinized soils can be excessively wet making establishment of saline tolerant vegetation difficult.

Soil features

Soils associated with Sandy ES are in the Mollisol order. Predominantly, they are classified further as Pachic Haplustolls, Typic Haplustolls, Pachic Argiustolls, and Typic Argiustolls. Included in this ecological are Typic Natrustolls (Desart series) which are deeper than 20 inches to a claypan layer. These soils were developed under prairie vegetation. They formed in weathered sandstone residuum and associated alluvium, in coarse-loamy eolian deposits over till, or in outwash deposits.

The common features of soils in this site are the moderately coarse textures to a depth exceeding 20 inches (forms a ribbon <1 inch long) and a drainage class of well to somewhat excessive. The surface layer is most commonly fine sandy loam or sandy loam, but loam is allowable if <10 inches thick. The soils are very deep to moderately deep. Moderately deep soils formed over sandstone. Some very deep soils have medium or coarse textured soil materials at a depth >20 inches.

In the moderately coarse-textured materials, salinity and sodicity typically are none. However, in the included Natrustolls the E.C. may be as high as 2 dS/m above the claypan and ranges from 2 to 8 dS/m in the claypan; the SAR may be as high as 5 above the claypan and ranges from 13 to 25 in the claypan. Soil reaction is strongly acid to slightly alkaline (pH 5.1 to 7.8) in the surface soil. Where a layer of carbonate accumulation occurs, it my increase to moderately alkaline (pH 7.9 to 8.4). In the included Natrustolls, the claypan is strongly alkaline (pH 8.5 to 9.0). Calcium carbonate content is none to moderately low (CaCO3 <5%) to a depth exceeding 20 inches. Deeper in the soil profile, each of these soil chemical properties may increase somewhat – particularly where medium-textured, contrasting materials occur.

Sub-surface soil layers are non-restrictive to water movement and root penetration. The hazard of water erosion increases on slopes greater than about 15 percent.

Major soil series correlated to the Sandy site are Desart, Dooley, Flaxton, Lefor, Livona, Manning, Parshall, Tally, Vebar, and Yegen.

Access Web Soil Survey (https://websoilsurvey.sc.egov.usda.gov/App/WebSoilSurvey.aspx) for specific local soils information.

Table 4. Representative soil features

Animal community

Animal Community – Wildlife Interpretations

Landscape

The MLRA 54 landscape is characterized by old, moderately dissected rolling plains with areas of local badlands, buttes, and isolated hills. MLRA 54 is considered to have a continental climate with cold winters and hot summers, low humidity, light rainfall, and much sunshine. Extremes in temperature are common and characteristic of MLRA 54. This area supports natural mixed-grass prairie vegetation with prairie rose, leadplant, and patches of western snowberry interspersed throughout the area. Green ash, chokecherry, and buffaloberry occur in draws and narrow valleys, creating woody riparian corridors. Complex, intermingled ecological sites create diverse grass- and shrubland habitats interspersed with varying densities of linear, slope, depressional, and in-stream wetlands associated with headwater streams and tributaries to the Missouri River. These habitats provide critical life-cycle components for many wildlife species.

Historic Communities/Conditions within MLRA 54:

The northern mixed-grass prairie was a disturbance-driven ecosystem with fire, herbivory, and climate functioning as the primary ecological drivers, either singly or often in combination. Many species of grassland birds, small mammals, insects, reptiles, amphibians, and herds of roaming bison, elk, and pronghorn were historically among the inhabitants adapted to this semi-arid region. Roaming herbivores, as well as several small mammal and insect species, were the primary consumers linking the grassland resources to large predators such as the wolf, mountain lion, and grizzly bear, and smaller carnivores such as the coyote, bobcat, red fox, and raptors. The black-tailed prairie dog was once abundant and provided ecological services by manipulating the plant and soil community providing habitat for the black-footed ferret, burrowing owl, ferruginous hawk, mountain plover, swift fox, small mammals, and amphibians and reptiles. Extirpated species include free-ranging American bison, grizzly bear, gray wolf, black-footed ferret, mountain plover, and peregrine falcon (breeding). Extinct from the region is the Rocky Mountain locust.

Present Communities/Conditions within MLRA:

Following European influence, domestic livestock grazing, elimination of fire, energy development, and other anthropogenic factors influenced plant community composition and abundance. Agriculture, transportation corridors, and energy development are the main factors contributing to habitat fragmentation, reducing habitat quality for area-sensitive species. These influences fragmented the landscape, reduced, or eliminated ecological drivers (fire), and introduced exotic species including smooth brome, crested wheatgrass, Kentucky bluegrass, and leafy spurge, further impacting plant and animal communities. The loss of the bison, black-tailed prairie dogs, and fire as primary ecological drivers greatly influenced the character of the remaining native plant communities and the associated wildlife moving towards a less diverse and more homogeneous landscape.

Included in this MLRA are the isolated Killdeer Mountains containing bur oak, quaking aspen, green ash, paper birch, western black birch, and American elm. Except for floodplain forests within the MRLA, the Killdeer Mountains contain the largest deciduous forest in southwestern North Dakota.

Some wildlife species in this area are mule deer, white-tailed deer, elk, pronghorn, coyote, red fox, bobcat, prairie rattlesnake, American badger, raccoon, North American porcupine, beaver, striped skunk, American mink, white-tailed jackrabbit, black-tailed prairie dog, Eastern and Merriam’s turkey, golden eagle, ferruginous hawks, sharp-tailed grouse, greater prairie chicken, black-billed magpie, and numerous species of grassland-nesting birds.

Presence of wildlife species is often determined by ecological site characteristics including grass and forb species, hydrology, aspect, and other associated ecological sites. The majority of species’ home range are larger than one ecological site or are dependent upon more than one ecological site for annual life requisites. Ecological sites offer different habitat elements as the annual life requisites change. Habitat improvement and creation must be conducted within the mobility limits of a known population for the species.

Insects play an important role providing ecological services for plant community development. Scavenging and decomposing insects provide the food chain baseline sustaining the carnivorous insects feeding upon them. Many insects provide the ecological services necessary for pollination, keeping plant communities healthy and productive. Insects provide a protein food source for numerous species including grassland-nesting birds and their young.

Species unique to the MLRA:

Bald Eagle: Bald Eagles prefer large rivers, lakes, reservoirs, or wetlands that are bordered by mature stands of trees or a single large tree. Bald eagles use the Missouri River system, including Lake Sakakawea, Lake Oahe, and associated tributaries. Mature trees, including cottonwoods, provide nesting sites adjacent to aquatic and upland foraging sites.

Dakota skipper: The extreme northern portion of this MLRA provides limited Dakota skipper habitat. Dakota skipper habitat is considered Type B habitat. Type B habitat is described as rolling native-prairie terrain over gravelly glacial moraine deposits dominated by bluestems and needlegrasses with the likely presence of bluebell bellflower, wood lily, purple coneflower, upright prairie coneflower, and blanket flower. The United States Fish and Wildlife Service lists two critical habitat units within the MLRA in McKenzie County, North Dakota.

Golden eagle: The Lake Sakakawea breaks, within the northwest portion of the MRLA 54, are key areas for golden eagle nesting. Grasslands, shrublands, and black-tailed prairie dog towns are used for foraging.

Black-footed ferret: Black-footed ferrets have been re-introduced as an experimental population in the southern portion of the MLRA located on the Cheyenne Sioux Indian Reservation. Since re-introduction between1991 and 1996, black-footed ferrets have been documented on the Standing Rock Sioux Indian Reservation approximately 20 miles north of the re-introduction site. Black-footed ferrets rely exclusively on prairie dog towns for shelter, breeding, and food sources (prairie dogs and other species within the town).

Least tern (Interior): Least terns are found on the Missouri River system in MLRA 54. Sparsely vegetated sandbars within the free-flowing portions of the Missouri River or shorelines of Lake Oahe and Sakakawea are used for nesting and foraging.

Species of Concern within the MLRA:

Following is a list of species considered “species of conservation priority” in the North Dakota State Wildlife Action Plan (2015); “species of greatest conservation need” in the Montana State Wildlife Action Plan (2015) and South Dakota State Wildlife Action Plan; and “species listed as threatened, endangered or petitioned” under the Endangered Species Act within MLRA 54 at the time this section was developed:

Invertebrates: Dakota skipper, little white tiger beetle, monarch butterfly, Ottoe skipper, regal fritillary, yellow-banded bumble bee, and western bumble bee.

Birds: American Kestrel, Baird’s sparrow, bald eagle, black-billed cuckoo, black tern, bobolink, Brewer’s sparrow, burrowing owl, chestnut-collared longspur, ferruginous hawk, golden eagle, grasshopper sparrow, greater sage-grouse, greater prairie-chicken, lark bunting, loggerhead shrike, least tern, long-billed curlew, marbled godwit, McCown’s longspur, mountain plover, northern goshawk, northern harrier, northern pintail, peregrine falcon (migration), piping plover, prairie falcon, red knot (migration), red-headed woodpecker, sharp-tailed grouse, short-eared owl, Sprague’s pipit, Swainson’s hawk, trumpeter swan, upland sandpiper, western meadowlark, willet, Wilson’s phalarope, and whooping crane (migration).

Mammals: Big and little brown bats, long-eared bat, long-legged bat, northern long-eared bat, Townsend’s big-eared bat, western small-footed bat, black-footed ferret, black-tailed prairie dog, dwarf shrew, gray wolf, hispid pocket mouse, Merriam’s shrew, northwestern moose, sagebrush vole, silver-haired bat, and swift fox.

Amphibians and Reptiles: Common snapping turtle, Great Plains toad, false map turtle, greater short-horned lizard, milk snake, northern leopard frog, plains hognose snake, plains spadefoot, sagebrush lizard, smooth green snake, and smooth softshell and spiny softshell turtle.

Fish and Mussels: Blue sucker, burbot, flathead chub, fragile papershell, northern redbelly dace, paddlefish, pallid sturgeon, pearl dace, pink papershell, shortnose gar, sickle-fin chub, sturgeon chub, and sauger.

Grassland Management for Wildlife in the MLRA

Management activities within the community phase pathways impact wildlife. Community phase, transitional, and restoration pathways are keys to long-term management within each State and between States. Significant inputs must occur to cross the threshold between States (e.g. State 3.0 to 2.0) requiring substantial economic inputs and management (mechanical, reseeding, prescribed fire, woody vegetation removal, grazing intensity, etc.). Timing, intensity, and frequency of these inputs can have dramatic positive or negative effects on local wildlife species. Ranchers and other land managers must always consider the long-term beneficial effects of management on the habitat in comparison to potential short-term negative effects to individuals.

Ecological sites occur as intermingled complexes on the landscape with gradual or sometimes abrupt transitions. Rarely do ecological sites exist in large enough acreage to manage independently. Ecological sites, supporting a dominance of herbaceous vegetation (Loamy/Limy Residual), can be located adjacent to ecological sites that support medium to tall shrubs (Loamy Overflow or Shallow). Conversely, ecological sites that are dominated by short- to mid-statured grasses (Claypan) can be adjacent to sites with bare soil only supporting a minor number of short grasses and forbs (Thin Claypan).

Management of these complex ecological sites can provide a heterogeneous or a homogenous landscape. Grassland bird use reduces as the plant community transitions to a homogenous state. Managers must recognize ecological sites and the complexes they occur in to properly manage the landscape. A management regime for one ecological site may negatively impact an adjacent site e.g., alteration of a grazing regime within a Flat Bottom Wooded Draw ecological site to encourage understory growth may encourage exotic cool-season grasses to increase or dominate an adjacent ecological site.

Life requisites and habitat deficiencies are determined for targeted species. Deficiencies need to be addressed along community phase, transitional, and restoration pathways as presented in specific state and transition models. Ecological sites should be managed and restored within the site’s capabilities to provide sustainable habitat. Managers also need to consider habitat provided by adjacent/intermingled ecological sites for species with home ranges or life requests that cannot be provided by one ecological site.

With populations of many grassland-nesting birds in decline, it is important to maintain these ecological sites in a 1.0 Reference State or the 2.0 Native/Invaded State. Plant communities, optimal for a guild of grassland species, serve as a population source where the birth rate exceeds mortality. Species may use marginal plant communities; however, these sites may function as a population sink where mortality exceeds the birth rate.

Understanding preferred vegetative stature and sensitivity to woody encroachment is necessary to manage for the specific grassland species. Various grass heights may be used for breeding, nesting, foraging, or winter habitat. While most species use varying heights, many have a preferred vegetative stature height. The following chart provides preferred vegetative stature heights and sensitivity to woody vegetation encroachment.

To view the chart, click on the hyperlink:
https://efotg.sc.egov.usda.gov/references/public/ND/54_Sandy_Narrative_FINAL_Ref_FSG.pdf

Sandy Wildlife Habitat Interpretation:

Sandy Ecological Sites are droughty sites identified by the presence of coarse sand. Sandy sites support drought-tolerant species but are still productive compared to loamy sites due to the increase in tall warm-season grasses like prairie sandreed. Sandy sites support diverse stands of tall and short warm-season grasses, along with diverse stands of cool-season grasses and numerous forb species. Associated ecological sites include sands, Shallow Sandy, Sandy Terrace, Sandy Claypan, and Loamy Overflow. This complex of ecological sites provides habitat for many edge-sensitive grassland bird species.

Sandy habitat features and components commonly support grassland-nesting birds, notably sharp-tailed grouse and greater prairie chicken nesting, brood cover, and lekking sites, depending upon its plant community state. Insects rely on associated forbs and grasses for survival, serve as food sources for birds and their young, and are forage for small and large herbivores.

Sandy Ecological Sites may be found in four plant community states (1.0 Reference State, 2.0 Native/Invaded State, 3.0 Short Grass State, 4.0 Invaded State, and 5.0 Go-Back State) within a local landscape. Multiple plant community phases exist within each state. Today, these states occur primarily in response to grazing and drought. Secondary influences include anthropogenic disturbances, black-tailed prairie dogs, and fire.

Because there is no known restoration pathway from State 2.0 to State 1.0, it is important to intensively manage using tools in the community phase pathways of State 1.0 and State 2.0 to prevent further plant community degradation along either the T1A Transitional Pathway to Native/Invaded State 2.0 or T2A Transitional Pathway to Short Grass State 3.0 thresholds. Native wildlife generally benefits from the heterogeneous grasslands, in stature and plant composition, found in Community Phases 1.1, 1.2, 2.1, and 2.2 which include diverse grass and forb species with varying stature and density. As plant communities degrade within State 2.0, short warm-season grasses, increase while native forbs are reduced. This transition results in reduced stature and increased plant community homogeneity. When adjacent/intermingled, ecological sites undergo the same transition, the result can be an expansive, homogenous landscape.

State 3.0 has a dramatic increase of short, warm-season grasses and club moss with a further reduction in native forbs. Reduced forb diversity limits insect populations, negatively affecting grassland-nesting bird foraging opportunities. Increased club moss can limit access to bare ground by nesting. A homogenous grassland landscape does not provide quality escape or winter cover. As a result, many species are not able to meet life requisites. Further degradation within State 3.0 occurs when cool-season exotic crested wheatgrass begins to dominate the site. Successful restoration along the R3A Restoration Pathway with native vegetation seeding and mechanical treatment will be expensive and dependent upon climatic and seedbed conditions.

Likewise, success along Restoration Pathway R4A from State 4.0 to State 2.0 is very difficult and is dependent upon presence of a remnant native grass population. This concept also applies to wildlife, as the target species must either be present on adjacent State 1.0 or State 2.0 plant communities, or on ecological sites within the species’ mobility limits. Species with limited mobility, such as Dakota skippers, must exist near the plant community in order to utilize restored sites. Mobile species such as grassland-nesting birds can easily locate isolated, restored plant communities.

Management along community phase, transition, or restoration pathways should focus on attainable changes. Short- and long-term monetary costs must be evaluated against short- and long-term ecological services in creating and maintaining habitat of sufficient quality to support a sustainable population density.

1.0 Reference State

Community Phase 1.1: Prairie Sandreed-Needlegrasses: This plant community offers good wildlife habitat, and every effort should be made to maintain this ecological site within this community phase. This phase retains high functionality through continued maintenance, including prescribed grazing with adequate recovery period as well as prescribed fire. Predominance of grass species in this community favors grazers and mixed-feeders (animals selecting grasses as well as forbs and shrubs). The structural diversity provides habitat for a wide array of migratory and resident birds.

Invertebrates: Insects play a role in maintaining the forb community and provide a forage base for grassland birds, reptiles, and rodents. Ecological services historically provided by bison are mirrored by domestic livestock. These services include putting plant material and dung in contact with mineral soil to be used by low trophic level consumers such as invertebrate decomposers, scavengers, shredders, predators, herbivores, dung beetles, and fungal-feeders.

Dakota skippers do not prefer this site due to limited host plants, such as little bluestem and prairie dropseed. Regal fritillary habitat is limited due to Nuttall's violet and prairie violets being uncommon. Monarch butterfly may use flowering forbs on this site; however, few milkweed species are found on this site to support breeding. The Sandy Ecological Site does not provide habitat for the little white tiger beetles, which prefer the large, active, Choppy Sands Ecological Site or sand beaches. This plant community provides habitat for the Ottoe Skipper, which prefers mid-to tall-statured grasses. Bumblebees and other native bees utilize forbs and bare ground for nesting amongst bunchgrasses. Prescribed grazing with adequate recovery periods, as well as prescribed fire to maintain the 1.1 Phase has little effect on nests of ground-dwelling insects.

Birds: This plant community provides quality nesting, foraging, and escape habitats favored by mid- to tall-grass nesting birds. Several species of grassland birds which prefer mid- to tall-grass stature s will use this site. In years with reduced precipitation or drought, nesting recruitment may be compromised. This plant community provides suitable areas for sharp-tailed grouse and greater prairie chicken nesting and brood-rearing habitat. Diverse prey populations provide good hunting opportunities for grassland raptors.

Mammals: The diversity of grasses and forbs provide high nutrition levels for small and large herbivores including voles, mice, rodents, jackrabbits, pronghorn, and white-tailed and mule deer. Short- to moderately-statured grasses provide suitable food and thermal, protective, and escape cover for small herbivores. The composition of mid- to tall-grasses may limit use by the hispid pocket mouse.

Amphibians and Reptiles: This ecological site and associated plant communities provide habitat for smooth green snakes. This ecological site can provide habitat for the northern leopard frog and Great Plains toad if freshwater habitat such as stock water ponds are located in or adjacent to the site. Plains spadefoot utilize small ephemeral ponds for breeding as well as other habitats offered within the Sandy ecological site. Sandy soils provide burrowing sites for short-horned lizards. Silver sagebrush and sandy soils provide excellent habitat for the sagebrush lizard.

Fish and Mussels: This ecological site is not directly associated with streams, rivers, or water bodies. Associated ecological sites, such as loamy overflow, can receive run-on hydrology from sandy sites. Management on these interconnected sites will have limited, secondary effects on aquatic species.

Community Phase 1.2 Needle and Thread–Prairie Sandreed/Sedge: Long-term, heavy, continuous grazing, repeated spring and summer drought, and/or annual early spring seasonal grazing increase the percentage of sedges in this plant community. This plant community becomes dominated by cool-season grasses, changing the stature of plant community from mid- to tall plant species to mid- to shortgrass species.

Invertebrates: Provides similar life requisites as Community Phase 1.1; however, heavy, continuous seasonal grazing may negatively impact ground-nesting sites for bumble bees, other native bees, and other ground-nesting insects due to reduction of forbs, timing of forb flowering, or increased soil compaction.

Birds: This plant community provides nesting, foraging, and escape habitats favored by short- to mid-grass nesting birds. A shift to shorter herbaceous plant stature along Community Phase Pathway 1.1A begins to benefit McCown’s longspur, chestnut-collared longspur, horned lark, and burrowing owl. Species preferring midgrass stature will be generally successful with normal to above normal precipitation and a change in management along the 1.2A Community Phase Pathway. In years with reduced precipitation or heavy grazing, nesting recruitment may be compromised for midgrass nesting species. This plant community provides areas suitable for sharp-tailed grouse and greater prairie chicken leks. Limited cover and diverse prey populations provide good hunting opportunity for grassland raptors.

Mammals: Provides similar life requisites as Community Phase 1.1.

Amphibians and Reptiles: Provides similar life requisites as Community Phase 1.1.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

Community Phase 1.3 Blue Grama/Sedge/Western Wheatgrass/Forbs: Short-statured grasses will dominate with continued heavy grazing, repeated drought, short-term prairie dog occupation, or a combination of these disturbances. Perennial grasses and forbs are reduced in stature and abundance.

Invertebrates: Reduction in forbs (stature and abundance) will reduce nectar availability for foraging insect populations. Continued heavy grazing may negatively impact ground nesting sites for bumble bees, other native bees, and other ground-nesting insects due to reduction of forbs, timing of forb flowering, or increased soil compaction. However, prairie dog burrows will provide nesting sites for bumble bees.

Birds: This plant community provides quality nesting, foraging, and escape habitats favored by shortgrass-nesting birds. A shift to shorter plant stature, along Community Phase Pathway 1.2B, benefits McCown’s longspur, chestnut-collared longspur, horned lark, and burrowing owl. Species preferring mid-grass stature may be successful with normal to above normal precipitation and a change in management along the 1.3A Community Phase Pathway. In years with reduced precipitation or heavy grazing, nesting recruitment will be compromised for mid-grass nesting species Limited cover and diverse prey populations provide good hunting opportunity for grassland raptors.

Mammals: Provides similar life requisites as Community Phase 1.1.

Amphibians and Reptiles: Provides similar life requisites as Community Phase 1.1.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

Community Phase 1.4 Annual Forbs/Bare Ground/Blue Grama/Sedges (Prairie Dog Town): This plant community phase is characterized by grazing-tolerant species and annual forbs. Prairie dog occupation will shift the plant community to increased annual forbs and grazing tolerant perennial grasses. Perennial forbs stature and abundance are being replaced by short-statured annual forbs. Bare ground increases while litter amounts and infiltration rates decline as soil surface temperatures increase. This short-statured plant community is resilient, retaining sufficient grazing-sensitive native species to return to 1.3 Community Phases (via Community Phase Pathway 1.4A).

Invertebrates: A switch to annual forbs from perennial forbs may not have a significant impact to invertebrates but may reduce season-long nectar producing plants for pollinators. Season-long nectar sources may be found on adjacent plant communities or ecological sites for mobile species. Increased bare ground and prairie dog burrow sites provide increased nesting sites for bumble bees and other ground-nesting insects.

Birds: This very short-statured phase, driven by continued over-grazing or prairie dog occupation, is favored by burrowing owls, chestnut-collared longspur, and McCown’s longspur. Prairie dog towns provide abundant prey populations for grassland raptors. The lack of grass and forb stature limits use by many bird species. Managing this phase along Community Phase Pathway 1.3A can be an economical and successful method to restore high quality habitat for many grassland-nesting birds.

Mammals: Suitable food, thermal, shelter, and escape cover (reduction in litter) for most mammals becomes limited. The loss of diversity of grasses and forbs reduces nutrition levels for small and large herbivores including rodents, white-tailed jackrabbits, and deer. Grazers, such as pronghorn, use prairie dog towns for foraging and loafing. Managing this phase along Community Phase Pathway 1.3A can be an economical and successful method to restore habitat.

Amphibians/Reptiles: Prairie dog towns provide habitat for both amphibians and reptiles. Tiger salamanders, prairie rattlesnakes, and other snake species will use the burrow systems of prairie dogs for shelter and denning.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

2.0 Native Invaded State

Community Phase 2.1 Prairie Sandreed-Needlegrasses-Western Wheatgrass: This plant community develops through Transition Pathway T1A, due to changes in management (chronic season-long or heavy late season grazing or complete rest) and the presence of exotic, cool-season grasses. The threshold between States 1.0 and 2.0 is crossed when Kentucky bluegrass, crested wheatgrass, smooth brome grass, or other exotic species become established. This plant community phase has a very similar appearance and function to the Reference State of Community 1.1, except it has a minor amount of cool-season exotic grasses and forbs. This phase functions at a high level for native wildlife, therefore, managers should consider the 2.0 Community Phase Pathways to avoid transitioning to State 3.0.

Invertebrates: Provides similar life requisites as Community Phase1.1.

Birds: Provides similar life requisites as Community Phase 1.1.

Mammals: Provides similar life requisites as Community Phase 1.1.

Amphibians and Reptiles: Provides similar life requisites as Community Phase1.1.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

Community Phase 2.2 Needle and Thread-Blue Grama/Sedge/Forbs: Continuous, heavy season-long grazing or heavy seasonal grazing along Community Phase Pathway 2.1A, leads to shorter-statured grasses, such as blue grama and sedges. Dominated by shorter-stature grasses and a loss of nitrogen-fixing or leguminous native forbs, the diversity of this plant community is reduced. Both tap-rooted and fibrous-rooted perennial forbs increase in this phase, but still remain a minor component. Prescribed grazing with adequate recovery periods along Community Phase Pathway 2.2A is an efficient, effective method to regain the cool-season grass and forb diversity components in Community Phase 2.1.

Invertebrates: The loss of native forbs and increase in sod-forming grasses limit foraging and nesting sites for all pollinators. Continuous, heavy season-long grazing or heavy seasonal grazing may reduce ground-nesting site availability. Homogeneity of forb species may limit season-long nectar availability.

Birds: Continuous, heavy season-long grazing or heavy seasonal grazing will reduce nesting sites, forage (invertebrates), and cover. A reduced forb component may limit foraging opportunities. Stature is generally short, serving both mid- and shortgrass-nesting birds. Shortgrass-nesting birds favor this phase. Species preferring midgrass stature will be generally successful with normal to above normal precipitation and a change in management along the 2.2A Community Phase Pathway. In years with reduced precipitation or heavy grazing during the nesting season, nesting recruitment may be compromised for species that prefer midgrass nesting. This plant community provides areas suitable for sharp-tailed grouse and greater prairie chicken lek site development. Limited stature and diverse prey populations provide good hunting opportunity for grassland raptors.

Mammals: Suitable food and thermal, protective, and escape cover (reduction in litter) for most mammals remains.

Amphibians and Reptiles: Provides similar life requisites as Community Phase 1.1.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

Community Phase 2.3 Blue Grama/Sedges/Western Wheatgrass/Forbs: Heavy, continuous grazing, with multiyear drought (along community phase pathway 2.2B) leads to an increase in blue grama and sedges; also, an increase in forbs (such as common yarrow, western rockjasmine, field chickweed, pussytoes, spiny phlox, and rough false pennyroyal) can be common along with shrubs such as prairie sagewort. The increase in sedges provides for a shorter statured plant community. Prescribed grazing with adequate recovery periods between grazing along with a return to average precipitation will shift the competitive edge to mid-statured native grasses species. This is the most effective method to regain diverse cool-season grass and forb components in community phase 2.2. Every effort should be made to manage within community phase pathway 2.3A to avoid crossing the threshold into State 3.0.

Insects: The increases in blue grama and sedges and a decrease in forb diversity limits foraging and nesting sites for all pollinators. Homogeneity of forb species may limit season long nectar availability.

Birds: Increases in blue grama and sedges results in a short-statured plant community. Native grasses are still present in the plant community; however, the increase in cool-season exotic grasses reduces plant structure favoring bird species that prefer mid- to short-grass species. Sharp-tailed grouse may use this plant community for lek sites; however, brood-rearing and winter cover may need to be provided by adjacent ecological sites or plant communities. Management for bird species preferring mid- to tall-statured grasses should follow community phase pathway 2.3A.

Mammals: The loss of diversity of grasses and forbs reduces nutrition levels for small and large herbivores. Short statured grasses and sedges limit thermal, protective, and escape cover for most mammals.

Amphibians/Reptiles: Provides similar life requisites as Community Phase 1.3.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

Community Phase 2.4 Annual Forbs/Bare Ground/Blue Grama/Sedges (Prairie Dog Town): This plant community is a result of ecological services provided by long-term black-tailed prairie dog occupation coupled with the introduction of exotic cool-season grasses and annual forbs (along Community Phase Pathway 2.3B or 2.2C). Black-tailed prairie dogs provide primary ecological services to transition to and maintain Plant Community Phase 2.4. Utilizing one or more tools in Community Phase Pathway 2.4A (e.g., removal of black- tailed prairie dogs, control of exotic perennial forbs, implementation of prescribed grazing) can move this community back to Phase 2.3, but this may require significant management and economic inputs.

Invertebrates: The loss of native forb diversity limits use by all pollinators. However, invasive forbs may provide limited seasonal use, dependent on bloom period. Bare ground, burrows, and short plant stature provide nest sites for bumblebees and other ground-nesting insects. Burrowing owls place dung around their burrow entrance, attracting dung beetles and other insects as a food source.

Birds: Burrowing owl and McCown’s longspur rely on the stature and composition that this plant community provides. Presence of black-tailed prairie dogs provides diverse prey populations for grassland raptors including burrowing owls, prairie falcons, and ferruginous hawks. Burrowing owls nest in abandoned prairie dog burrows.

Mammals: Suitable food, thermal, protective, and escape cover (reduction in litter) for most mammals becomes limited. The loss of grass and forb diversity reduces nutrition levels for small and large herbivores including voles, mice, rodents, white-tailed jackrabbits, cottontail rabbits, and deer. Except for black-tailed prairie dog, this plant community provides little habitat for mid-sized or small herbivores. Nonetheless, black-tailed prairie dog towns provide important habitat for many mammal species including small rodents. Grazers, such as pronghorn, use prairie dog towns for foraging and loafing.

Amphibians/Reptiles: Prairie dog towns provide habitat for both amphibians and reptiles. Tiger salamanders, prairie rattlesnakes, and other snake species will use the burrow systems of prairie dogs for shelter and denning.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

3.0 Short Grass State

through community phase pathway T2A, results in a plant community dominated by blue grama and lesser spikemoss. Lesser spikemoss can cover 25% or greater of the area. Perennial forbs have been replaced by annual forbs with bloom periods variable dependent on climatic conditions. Once lesser spikemoss has become well established, mechanical (as well as other) treatments via Restoration Pathway R3A have been largely ineffective in suppressing or eliminating it, even though some short-term reductions may appear successful. As a result, this restoration pathway will likely be temporary.

Insects: Lesser spikemoss does not provide any nectaring opportunities while blue grama can form a sod-like surface, reducing or eliminating ground nesting bee opportunities. The loss of native forbs and dominance of lesser spikemoss limit foraging and ground nesting sites for pollinating insects.

Birds: Blue grama/lesser spikemoss benefits species dependent on short grass structure. A reduced forb component may limit foraging opportunities. Sharp-tailed grouse may still use this plant community for leks where lesser spikemoss does not dominate. Management for bird species preferring mid- to tall-statured grasses should follow community phase pathway T2A.

Mammals: Shorter statured grasses reduce thermal cover and protection for mammals. Foraging become limited as lesser spikemoss provide limited foraging opportunities.
Amphibians/Reptiles: Blue grama and lesser spikemoss limit bare ground which negatively impacts burrowing opportunities for the short-horned lizard.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

Community Phase 3.2 Crested Wheatgrass/Blue Grama/Lesser Spikemoss: Failed mechanical treatment of lesser spikemoss allows cool-season exotic grass, such as crested wheatgrass, to occupy this site.

Invertebrates: Provides similar life requisites as community phase 3.1.

Birds: Crested wheatgrass and blue grama benefit species dependent on short grass structure. A reduced forb component may limit foraging opportunities. Sharp-tailed grouse may still use this plant community for leks where lesser spikemoss does not dominate. However, brood-rearing habitat is somewhat better than 3.1 with created wheatgrass providing limited brood cover; however, the loss of native forbs may limit foraging opportunities for chicks. Management for bird species preferring mid- to tall-statured grasses should follow community phase pathway T2A.

Mammals: Provides similar life requisites as community phase 2.2. Amphibians/Reptiles: Provides similar life requisites as community phase 3.1. Fish and Mussels: Provides similar life requisites as community phase 1.1.

4.0 Invaded State

Community Phase 4.1 Exotic Cool-Season Grasses/Shrubs: Community phase pathway T2B is characterized by non-use (10 or more years) or low intensity (<20% utilization) grazing and elimination of fire when exotic cool-season grasses are present, as in community phase 2.0. Community phase pathway T3A will also lead to 4.1. Native forbs continue to decrease in abundance and shrubs, such as rose and western snowberry, tend to increase. Restoration pathway R4A requires remnant amounts of native warm-season grasses (i.e., blue grama, prairie sandreed), cool-season grasses (i.e., needlegrasses, western wheatgrass), and forbs. These remnant populations can only be expressed through frequent prescribed burns and high levels of prescribed grazing management targeting the exotic cool-season grasses. Intensified management along the R4A pathway may have significant short-term negative impacts on wildlife habitat; however, this is necessary to restore long term native habitat functions.

Invertebrates: Non-use or low intensity (<20% utilization) grazing limits use by beneficial insects provided in States 1.0 and 2.0. Increased litter and lack of grazing lead to limited contact between plant material and mineral soil resulting in a cooler micro-climate, which is unfavorable to most insects. Lack of bare soil limits ground nesting sites for native bees and other ground nesting insects. The lack of nectar producing plants limits forage opportunities for bumblebees, regal fritillary, monarch butterfly, and other pollinating species.

Birds: This homogeneous community phase, dominated by exotic plant species, provides limited habitat and life requisites for most obligate grassland nesting birds. Lack of structure and plant diversity, along with increased litter and the tendency of Kentucky bluegrass and smooth brome to lay down, limits use by many grassland nesting birds. Litter accumulations reduce use by chestnut-collared and McCown’s longspurs. Western snowberry reduces use of this site by species that avoid areas with woody vegetation. Sharp-tailed grouse may use these sites for lekking but provide limited brood rearing and winter cover.

Mammals: Litter accumulation favors thermal, protective, and escape cover for small rodents. However, reduced availability of native grass seed may reduce food availability for species such as the hispid pocket mouse.

Amphibians/Reptiles: Provides similar life requisites as community phase 1.1. However, increased litter and cooler soil temperature may reduce use by plains spadefoot and short-horned lizard.

Fish and Mussels: Provides similar life requisites as community phase 1.1.

5.0 Go-Back State

Community Phase 5.1 Annual/Pioneer Perennial/Exotics: These plant communities are the result of severe soil disturbance (such as cropping, recreational activity, or concentrated livestock activity for a prolonged period). Following cessation of disturbances, the resulting plant community is dominated by early pioneer annual and perennial plant species. Plant species composition and production are highly variable. Weedy plants can provide pollinator habitat along with spring and summer cover for many mammals and birds, and their young. Dense weed cover can keep soils moist, increasing insect presence. Tall structure provided by some weeds, such as marsh elder and ragweed, offer thermal cover and seeds throughout winter.

Successful restoration of native species along transition pathway R5A results in a native grass and forb community in State 2.0. Failed restoration to native species through restoration pathway R5B results in Invaded State 3.0. Wildlife species response will be dependent upon plant community composition, vegetative structure, patch size, and management activities (such as prescribed grazing, burning, interseeding, haying, or noxious weed control).

Animal Community – Grazing Interpretations

This site is well adapted to managed grazing by domestic livestock. The predominance of herbaceous plants across all plant community phases best lends these sites to grazing by cattle, but other domestic grazers with differing diet preferences may also be a consideration depending upon management objectives. Often, the current plant community does not match any particular plant community (as described in the ecological site description). Because of this, a resource inventory is necessary to document plant composition and production. Proper interpretation of this inventory data will permit the establishment of a safe, initial stocking rate for the type and class of animals and level of grazing management. More accurate stocking rate estimates should eventually be calculated using actual stocking rate information and monitoring data.

NRCS defines prescribed grazing as “managing the harvest of vegetation with grazing and/or browsing animals with the intent to achieve specific ecological, economic, and management objectives”. As used in this site description, the term ‘prescribed grazing’ is intended to include multiple grazing management systems (e.g., rotational grazing, twice-over grazing, conservation grazing, targeted grazing, etc.) provided that, whatever management system is implemented, it meets the intent of prescribed grazing definition.

The basic grazing prescription addresses balancing forage demand (quality and quantity) with available forage, varying grazing and deferment periods from year-to-year, matching recovery/deferment periods to growing conditions when pastures are grazed more than once in a growing season, implementation of a contingency (e.g., drought) plan, and a monitoring plan. When the management goal is to facilitate change from one plant community phase or state to another, then the prescription needs to be designed to shift the competitive advantage to favor the native grass and forb species.

Grazing levels are noted within the plant community narratives and pathways in reference to grazing management. “Degree of utilization” is defined as the proportion of the current years forage production that is consumed and/or destroyed by grazing animals (may refer to a single plant species or a portion or all the vegetation). “Grazing utilization” is classified as slight, moderate, full, close, and severe (see the following table for description of each grazing use category). The following utilization levels are also described in the Ranchers Guide to Grassland Management IV. Utilization levels are determined by using the landscape appearance method as outlined in the Interagency Technical Reference “Utilization Studies and Residual Measurements” 1734-3.

Utilization Level % Use Description
Slight (Light) 0-20 Appears practically undisturbed when viewed obliquely. Only choice areas and forage utilized.

Moderate 20-40 Almost all of accessible range shows grazing. Little or no use of poor forage. Little evidence of trailing to grazing.

Full 40-60 All fully accessible areas are grazed. The major sites have key forage species properly utilized (about half taken, half left). Points of concentration with overuse limited to 5 to 10 percent of accessible area.

Close (Heavy) 60-80 All accessible range plainly shows use and major sections closely cropped. Livestock forced to use less desirable forage, considering seasonal preference.

Severe > 80 Key forage species completely used. Low-value forages are dominant.

Hydrological functions

Available water is the principal factor limiting herbage production on this site. The site is dominated by soils in hydrologic group B with localized areas in groups A and C. Infiltration varies from moderately rapid to moderate; runoff potential varies from very low to high for this site depending on soil hydrologic group, surface texture, slope percent, slope shape, and ground cover. In many cases, areas with greater than 75% ground cover have the greatest potential for high infiltration and lower runoff. An exception would be where shortgrasses form a dense sod and dominate the site. Areas where ground cover is less than 50% have the greatest potential to have reduced infiltration and higher runoff (refer to Section 4, NRCS National Engineering Handbook for runoff quantities and hydrologic curves).

Recreational uses

The largest acreage of public land available for recreation in the MLRA is owned and managed by the United States Forest Service (USFS) within the Little Missouri, Grand River, and Cedar River National Grasslands in South Dakota and the Little Missouri National Grasslands in North Dakota (687,398 acres). These areas are available for hunting, fishing, hiking, camping, horse and bike riding, nature viewing, etc. In addition, the Bureau of Land Management (BLM) manages (40,264 acres) in North and South Dakota with the same recreational opportunities as the USFS lands. The United States Army Corps of Engineers (USAE) owns 496,162 acres of land and water located on and adjacent to Lake Sakakawea and Lake Oahe. The North Dakota and South Dakota Game and Fish Departments manage the fisheries resources. These two Missouri River reservoirs provide excellent fishing and water recreation opportunities. In addition, the United States Fish and Wildlife Service (USFWS) manages a national fish hatchery below Garrison Dam.

The USFWS manages 36,858 acres in the National Wildlife Refuge system while the North Dakota and South Dakota wildlife management agencies manage 72,218 acres as wildlife or game management areas. The North Dakota, South Dakota, and Montana Department of Trust Lands manage 486,482 acres. These areas provide hunting, bird watching, hiking, and other outdoor recreation opportunities. North Dakota Wildlife Management Areas along the shoreline of Lake Sakakawea and the Missouri River account for 60,000 acres of the approximately 72,218 acres of land managed by the states for wildlife habitat in MLRA 54. Located in the northern portion of the MLRA, the Killdeer Mountain WMA is the largest tract of state-owned land managed for wildlife habitat at approximately 7,000 acres.

The largest refuge managed by the United States Fish and Wildlife service is Lake Ilo National Wildlife Refuge totaling approximately 4,000 acres. United States Bureau of Reclamation manages approximately 11,000 acres at Lake Tschida and 8,460 acres at Bowman-Haley Lake for fish and wildlife habitat. The National Park Service manages the Knife River Indian Village National Historic Site; the North Dakota Historical Society manages the Double Ditch Indian Village site.

Bird watching: Public and private grasslands within MLRA 54 provide essential habitat for prairie- dependent bird species such as Sprague's pipits, western meadowlark, and Baird's sparrow along with some of the larger, showy members of the upland prairie include marbled godwits, upland sandpipers, willets, and sharp-tailed grouse. Publicly owned lands provide excellent birding opportunities. MLRA 54 is in the Central Flyway.

Hunting/Fishing: MLRA 54 is a fall destination for thousands of pheasant and upland game bird hunters. This MLRA also provides excellent deer (white-tailed and mule), pronghorn, and coyote hunting opportunities. Lake Sakakawea, Lake Oahe, Lake Tschida, and the Missouri River provide excellent year-round fishing opportunities. The North Dakota Game and Fish Department and South Dakota Game, Fish and Parks manage approximately 40 fishing lakes within the MLRA. Available species include yellow perch, walleye, northern pike, muskellunge, crappie, bluegill, rainbow trout, and smallmouth bass. Chinook salmon are stocked in Lake Sakakawea.

Camping: Numerous state operated campgrounds are located along the shores of Lake Sakakawea, Lake Oahe, Missouri River, and Shadehill Reservoir. Primitive camping is allowed on Grand River and Cedar River National Grasslands in South Dakota and the Little Missouri National Grasslands in North Dakota. Other numerous camping (primitive and improved) sites are available in numerous city and county parks.

Hiking/Biking/Horseback Riding: Hiking is permitted on most state and federally owned lands. Developed hiking and biking trails can be found on Harmon Lake (13.1 miles), Roughrider Trail (Morton County, 16.5 miles), Missouri River State Natural Area (5 miles), Ft. Abraham Lincoln State Park (8 miles), Cross Ranch State Park (14 miles), Grand River National Grasslands (7 miles), Lake Sakakawea State Park (5 miles), and Lewis & Clark State Park (5 miles). In addition, extensive biking and walking trails are found in local county and city parks. Ft. Abraham Lincoln State Park has 6 miles of horseback trails.

Wood products

No appreciable wood products are present on the site.

Other products

Seed harvest of native plant species can provide additional income on this site.

Other information

Site Development and Testing Plan
• The range of slopes for this site is very wide (0 to 50% slopes). Future investigation of plant communities and forage production on steeper slopes is recommended.
• NASIS revisions needed:
o Saline phases of Vebar and Parshall are currently linked to the Saline Lowland ecological site. These soils are moderately well drained or well drained. These saline phases commonly developed due to cultivation (seeps). Range planting considerations need to include factors other than the presence of accumulated salts (recharge area, etc.). Vebar components need to be relinked to Sandy. Parshall components need to be relinked to Sandy or Loamy Overflow (as appropriate).
o Appam and Lehr are currently linked to the Sandy. These soils have sand and gravel within a depth of 20 inches and should be linked to a Shallow Gravel ESD. MLRA 54 does not have a Shallow Gravel ESD; these components should be linked to 53B Shallow Gravel.
o Some Parshall components need to be relinked from Sandy to Loamy Overflow.
This ESD is the best available knowledge. The site concept and species composition table have been used in the field and tested for more than 5 years. It is expected that as additional information becomes available revisions may be needed.

Inventory data references

Information presented here has been derived from NRCS and other federal/state agency clipping and inventory data. Also, field knowledge of range-trained personnel was used. All descriptions were peer reviewed and/or field-tested by various private, state and federal agency specialists.

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Contributors

ND NRCS: David Dewald, Jonathan Fettig, Jody Forman, Mike Gerbig, Alan Gulsvig, Mark Hayek, Jeanne Heilig, John Kempenich, Chuck Lura, Jeff Printz, Steve Sieler, and Hal Weiser.

Approval

Suzanne Mayne-Kinney, 3/31/2025

Acknowledgments

NRCS would like to acknowledge the United State Forest Service (USFS) and National Park Service (NPS) for access to USFS properties and technical assistance in ESD development. USFS: Jack Dahl, Nickole Dahl, and Chad Prosser.