Physiographic features

This site occurs on nearly level to very steep sedimentary plains; it occurs on hillslopes, pediments and alluvial flats. Parent materials are residuum or alluvium from residuum. Slope ranges from 0 to 45 percent.

Table 2. Representative physiographic features

Climatic features

MLRA 54 is considered to have a continental climate with cold winters and hot summers, low humidity, light rainfall, and much sunshine. Extremes in temperature are common and characteristic of the MLRA. The continental climate is the result of this MLRA’s location in the geographic center of North America. There are few natural barriers on the northern Great Plains, so air masses move unobstructed across the plains and account for rapid changes in temperature.

Annual precipitation ranges from 14 to 18 inches per year. The normal average annual temperature is about 42° F. January is the coldest month with average temperatures ranging from about 13° F (Beach, ND) to about 16° F (Bison, SD). July is the warmest month with temperatures averaging from about 69° F (Beach, ND) to about 72° F (Timber Lake, SD). The range of normal average monthly temperatures between the coldest and warmest months is about 57° F. This large temperature range attests to the continental nature of MLRA 54’s climate. Wind speeds average about 11 miles per hour, ranging from about 13 miles per hour during the spring to about 10 miles per hour during the summer. Daytime wind speeds are generally stronger than nighttime wind speeds, and occasional strong storms may bring brief periods of high winds with gusts to more than 50 miles per hour.

Growth of native cool-season plants begins in late March and continues through early to mid-July. Native warm-season plants begin growth in mid-May and continue to the end of August. Greening up of cool-season plants can occur again in September and October when adequate soil moisture is present.

Table 3. Representative climatic features

Climate stations used

• (1) FT YATES 4 SW [USC00323207], Fort Yates, ND

• (2) HETTINGER [USC00324178], Hettinger, ND

• (3) DUPREE [USC00392429], Dupree, SD

• (4) WATFORD CITY [USC00329233], Watford City, ND

• (5) MANDAN EXP STN [USC00325479], Mandan, ND

• (6) LUDLOW 3 SSE [USC00395048], Ludlow, SD

• (7) HEBRON [USC00324102], Hebron, ND

Influencing water features

No significant water features influence this site. A seasonal ground water table is typically deeper than 5 feet throughout the growing season. Soils occurring on fans may receive some run-on water from adjacent uplands, but this does significantly influence the plant community. Permeability is moderately slow to moderate. Water loss is through evapotranspiration and percolation below the root zone.

Salinization Risk – The potential salinization of the site can occur in any of the vegetative states; however, it is most prevalent in the State 4.0 Go-Back. Removal of perennial vegetation (e.g., annual cropping), above this ecological site (recharge area), results in increased water moving down through the soil profile. This downward water movement leaches salts over time creating shallow saline groundwater immediately above a less permeable layer. Due to gravity, water moves downward through the soil profile then laterally through a porous layer, such as a coal or sand/gravel seam, transporting salts to the discharge area. Below the porous layer, is a less permeable layer such as soft sedimentary bedrock. Where shallow saline groundwater occurs, salts often concentrate at or near the soil surface through capillary rise (discharge area). In capillary rise, water moves from where the soil is saturated, or nearly so, to drier soil against the force of gravity. Evaporation at the soil surface dries the soil and “pulls” water by capillary flow from the wet soil zone. Because only pure water evaporates, salts are left behind.

Figure 7. Increased salinity levels will make establishment of grasses and forbs more difficult. Treatment of the recharge area is a requirement to reclaim these soils from salinization. Some salinized soils can be excessively wet making establishment of saline tolerant vegetation difficult.

Soil features

Soils associated with the Limy Residual ES are in the Mollisol and Inceptisol orders. The Mollisols are classified further as Typic Calciustolls. The Inceptisols are classified further as Typic Haplustepts and Calcic Haplustepts. These soils were developed under prairie vegetation. They formed in calcareous, medium- textured residuum (typically siltstone) or slope alluvium eroding from the adjacent uplands. The soils on this site are moderately deep (>20 inches to soft, sedimentary bedrock) to very deep. The sedimentary beds affect root growth. The soils are well drained. Textural family classes are mostly fine silty but fine-loamy may also occur.

This ecological site occurs on two primary landforms: hillslopes and alluvial fans. The residual soils developed on the shoulders, back slopes, and foot slopes of gently sloping to moderately steep hillslopes and pediments. The alluvial soils formed in slope alluvium on relatively stable linear fans. These fans are well vegetated since deposition and erosion are occurring at relatively slower rates than the actively eroding depositional fans (Badlands Fan ecological site) directly adjacent to very steep, soft sedimentary bedrock, (e.g., Badlands). Thin, stratified layers may be evident in the parent material on the fan landform.

Characteristics common to the soil in this site are a dark grayish brown to yellowish brown surface layer (4 to 8 inches thick). Silt loam is the predominant texture above the sedimentary beds, but loam or silty clay loam also occur. The soil forms a ribbon 1 to 2 inches long. Depth to calcium carbonates is <8 inches; typically, the soil effervesces throughout.

Soil reaction typically is neutral to moderately alkaline (pH 6.6 to 8.4) in the surface layer and slightly alkaline to moderately alkaline (pH 7.4 to 8.4) in the subsoil. Salinity is generally none to very slight (E.C. 0 to 4 dS/m); however, in some soils it is slight (E.C. 4 to 8 dS/m) in the substratum. Sodicity is generally low (SAR <5); however, in some soils, the SAR is as high as 10 in the lower subsoil and substratum. Calcium carbonate content typically ranges from 5 to 40 percent. Some soils in native grassland have <5% CaCO3 in the surface few inches.

These soils are particularly susceptible to water and wind erosion. The hazard of water and wind erosion increases where vegetative cover is sparse or non-existent. Loss of the soil surface layer results in a shift in species composition and production.

The major soil series which characterize the Limy Residual ecological site are Chama, Cherry, Lantry, and Maschetah.

Access Web Soil Survey (https://websoilsurvey.sc.egov.usda.gov/App/WebSoilSurvey.aspx) for specific local soils information.

Table 4. Representative soil features

Animal community

Animal Community – Wildlife Interpretations

Landscape

The MLRA 54 landscape is characterized by moderately dissected rolling plains with areas of local badlands, buttes, and isolated hills. MLRA 54 is considered to have a continental climate with cold winters and hot summers, low humidity, light rainfall, and much sunshine. Extremes in temperature are common and characteristic of the MLRA. This area supports natural mixed-grass prairie vegetation with prairie rose, leadplant, and patches of western snowberry interspersed throughout the area. Green ash, chokecherry, and buffaloberry occur in draws and narrow valleys creating woody riparian corridors. Complex intermingled ecological sites create diverse grass/shrub land habitats interspersed with varying densities linear, slope, depressional, and in-stream wetlands associated with headwater streams and tributaries to the Missouri River. These habitats provide critical life-cycle components for many wildlife species.

Historic Communities/Conditions within MLRA:

The northern mixed-grass prairie was a disturbance-driven ecosystem with fire, herbivory, and climate functioning as the primary ecological drivers (either singly or often in combination). Many species of grassland birds, small mammals, insects, reptiles, amphibians, and large herds of roaming American bison, elk, and pronghorn were historically among the inhabitants adapted to this semi-arid region. Roaming herbivores, as well as several small mammal and insect species, were the primary consumers linking the grassland resources to large predators (such as the wolf, mountain lion, and grizzly bear) and smaller carnivores (such as the coyote, bobcat, red fox, and raptors). The black-tailed prairie dog was once abundant and provided ecological services by manipulating the plant and soil community, thus providing habitat for the black-footed ferret, burrowing owl, ferruginous hawk, mountain plover, swift fox, small mammals, and amphibians and reptiles. Extirpated species include free-ranging American bison, grizzly bear, gray wolf, black- footed ferret, mountain plover, and peregrine falcon (breeding). Extinct from the region is the Rocky Mountain locust.

Present Communities/Conditions within MLRA:

Following European influence, domestic livestock grazing, elimination of fire, energy development, and other anthropogenic factors influenced plant community composition and abundance. Agriculture, transportation corridors, and energy development are the main factors contributing to habitat fragmentation, reducing habitat quality for area-sensitive species. These influences fragmented the landscape; reduced or eliminated ecological drivers (fire); and introduced exotic species including smooth brome, crested wheatgrass, Kentucky bluegrass, and leafy spurge. This further impacted plant and animal communities. The loss of the bison, black-tailed prairie dogs, and fire as primary ecological drivers greatly influenced the character of the remaining native plant communities and the associated wildlife, moving towards a less diverse and more homogeneous landscape. Included in this MLRA are the isolated Killdeer Mountains (limestone capped residual butte) containing bur oak, quaking aspen, green ash, paper birch, and American elm. Except for floodplain forests within the MLRA, the Killdeer Mountains contain the largest deciduous forest in southwestern North Dakota.

Some wildlife species in this area are mule deer, white-tailed deer, elk, pronghorn, moose, coyote, red fox, bobcat, prairie rattlesnake, American badger, raccoon, North American porcupine, beaver, striped skunk, American mink, white-tailed jackrabbit, black-tailed prairie dog, Eastern and Merriam’s turkey, golden eagle, ferruginous hawks, sharp-tailed grouse, black-billed magpie, and numerous species of grassland-nesting birds and pollinating insects.

Presence of wildlife species is often determined by ecological site characteristics including grass and forb species, hydrology, aspect, and other associated ecological sites. The home ranges of a majority species are larger than one ecological site or are dependent on more than one ecological site for annual life requisites. Ecological sites offer different habitat elements as the annual life requisites change. Habitat improvement and creation must be conducted within the mobility limits of a known population for the species.

Insects play an important role providing ecological services for plant community development. Insects that are scavengers or aid in decomposition provide the food chain baseline sustaining the carnivorous insects feeding upon them. Many insects provide the ecological services necessary for pollination, keeping plant communities healthy and productive. Insects provide a protein food source for numerous species including grassland-nesting birds and their young.

Species unique to MLRA:

Bald eagle: Bald Eagles prefer large rivers, lakes, reservoirs, or wetlands that are bordered by mature stands of trees or a single large tree. Bald eagles use the Missouri River system, including Lakes Sakakawea and Oahe, and associated tributaries. Mature trees, including cottonwoods, provide nesting sites adjacent to aquatic and upland foraging sites.

Dakota skipper: The extreme northern portion of this MLRA provides limited Dakota skipper habitat. Dakota skipper habitat within MLRA 54 is considered Type B habitat. Type B habitat is described as rolling native-prairie terrain over gravelly glacial moraine deposits dominated by bluestems and needlegrasses with the likely presence of bluebell bellflower, wood lily, blacksamson echinacea, upright prairie coneflower, and blanketflower. The United States Fish and Wildlife Service lists two critical habitat units within the MLRA in McKenzie County, North Dakota.

Golden eagle: The Lake Sakakawea breaks, bluffs, and rock outcroppings within the northwest portion of the MLRA are key areas for golden eagle nesting. Grasslands, shrublands, and black-tailed prairie dog towns are used for foraging.

Black-footed ferret: Black-footed ferrets have been reintroduced as an experimental population in the southern portion of the MLRA located on the Cheyenne Sioux Indian Reservation. Since reintroduction between 1991 and 1996, black-footed ferrets have been documented on the Standing Rock Sioux Indian Reservation approximately 20 miles north of the reintroduction site. Black-footed ferrets rely exclusively on prairie dog towns for shelter, breeding, and food sources (prairie dogs and other species within the town).

Least tern (Interior): Least terns are found on the Missouri River system in MLRA 54. Sparsely vegetated sandbars within the free-flowing portions of the Missouri River or shorelines of Lake Oahe and Sakakawea are used for nesting and foraging.

Species of Concern within the MLRA:

The following is a list of species considered “species of conservation priority” in the North Dakota State Wildlife Action Plan (2015); “species of greatest conservation need” in the Montana State Wildlife Action Plan (2015) and the South Dakota State Wildlife Action Plan (2014); and “species listed as threatened, endangered, or petitioned” under the Endangered Species Act within MLRA 54 at the time this section was developed:

Invertebrates: Dakota skipper, little white tiger beetle, monarch butterfly, Ottoe skipper, regal fritillary, yellow-banded bumble bee, and western bumble bee.

Birds: American Kestrel, Baird’s sparrow, bald eagle, black-billed cuckoo, black tern, bobolink, Brewer’s sparrow, burrowing owl, chestnut-collared longspur, ferruginous hawk, golden eagle, grasshopper sparrow, greater sage-grouse, lark bunting, loggerhead shrike, least tern, long-billed curlew, marbled godwit, McCown’s longspur, mountain plover, northern goshawk, northern harrier, northern pintail, peregrine falcon (migration), piping plover, prairie falcon, red knot (migration), red- headed woodpecker, sharp-tailed grouse, short-eared owl, Sprague’s pipit, Swainson’s hawk, trumpeter swan, upland sandpiper, western meadowlark, willet, Wilson’s phalarope, and whooping crane (migration).

Mammals: Big and little brown bats, long-eared bat, long-legged bat, northern long-eared bat, Townsend’s big-eared bat, western small-footed bat, black-footed ferret, black-tailed prairie dog, dwarf shrew, gray wolf, hispid pocket mouse, Merriam’s shrew, northwestern moose, sagebrush vole, silver-haired bat, and swift fox.

Amphibians/Reptiles: Common snapping turtle, Great Plains toad, false map turtle, greater short- horned lizard, milk snake, northern leopard frog, plains hognose snake, plains spadefoot, smooth green snake, and smooth softshell and spiny softshell turtle.

Fish and Mussels: Blue sucker, burbot, flathead chub, fragile papershell, northern redbelly dace, paddlefish, pallid sturgeon, pearl dace, pink papershell, shortnose gar, sickle-fin chub, sturgeon chub, and sauger.

Grassland Management for Wildlife in the MLRA

Management activities within the community phase pathways impact wildlife. Community phase, transitional, and restoration pathways are keys to long-term management within each state and between states. Significant inputs must occur to cross the threshold between states (e.g., State 3.0 to 2.0) requiring substantial economic inputs and management (mechanical, reseeding, prescribed burning, woody vegetation removal, grazing intensity, etc.). Timing, intensity, and frequency of these inputs can have dramatic positive or negative effects on local wildlife species. Ranchers and other land managers must always consider the long-term beneficial effects of management on the habitat in comparison to potential short-term negative effects to individuals.

Ecological sites occur as intermingled complexes on the landscape with gradual or sometimes abrupt transitions. Rarely do ecological sites exist in large enough acreage to manage independently. Ecological sites that support a dominance of herbaceous vegetation (Loamy/Clayey) can be located adjacent to ecological sites that support medium to tall shrubs (Loamy Overflow). Conversely, ecological sites that are dominated by short- to mid-statured grasses (Claypan) can be adjacent to sites with bare soil only supporting minor amounts of short grasses and forbs (Thin Claypan).

Management of these complex ecological sites can provide a heterogeneous or a homogenous landscape. Grassland bird use declines as the plant community transitions to a homogenous state. Managers should recognize ecological sites and the complexes they occur in to properly manage the landscape. A management regime for one ecological site may negatively impact an adjacent site; for example, alteration of a grazing regime within a Loamy Overflow ecological site to encourage understory growth may encourage exotic cool-season grasses to increase or dominate an adjacent ecological site.

Life requisites and habitat deficiencies are determined for targeted species. Deficiencies must be addressed along community phase, transitional, and restoration pathways as presented in specific state-and-transition models. Ecological sites should be managed and restored within the site’s capabilities to provide sustainable habitat. Managers also should consider habitat provided by adjacent/intermingled ecological sites for species with home ranges or life requisites that cannot be provided by one ecological site.

With populations of many grassland-nesting birds in decline, it is important to maintain these ecological sites in a 1.0 Reference State (rarely found intact) or the 2.0 Native/Invaded. Plant communities optimal for a guild of grassland species serve as a population source where the birth rate exceeds mortality. Species may use marginal plant communities; however, these sites may function as a population sink where mortality exceeds the birth rate.

Understanding preferred vegetative stature and sensitivity to woody encroachment is necessary to manage for the specific grassland species. Various grass heights may be used for breeding, nesting, foraging, or winter habitat. While most species use varying heights, many have a preferred vegetative stature height. The following chart provides preferred vegetative stature heights and sensitivity to woody vegetation encroachment.

For more information:
https://efotg.sc.egov.usda.gov/references/public/ND/54_Limy_Residual_Narrative_FINAL_Ref_FSG.pdf

Limy Residual Wildlife Interpretation:

Limy Residual ecological sites are defined by calcium carbonates in the upper part of the soil profile (within a depth of 8 inches) and soil parent material that weathered from residuum or is alluvium from residuum. They are less productive and less diverse than Loamy and Loamy Overflow sites which occur on similar landscape positions. Limitations within Limy Residual sites support shorter stature and lower diversity of grasses and forbs for wildlife. The expansiveness of the Loamy, Loamy Terrace, Shallow Loamy, Very Shallow and Thin Loamy ecological sites, associated with the Limy Residual sites in MLRA 54, provides generally an unfragmented landscape. This complex of ecological sites provides habitat for many edge-sensitive grassland bird species. Also, in the Killdeer Mountains, the Upland Hardwood Forest ecological site is associated with the Limy Residual site.

Limy Residual vegetative features and components commonly support grassland-nesting birds. Notably, the shorter grass on Limy Residual sites can provide sharp-tailed grouse lekking sites. Insects rely on associated forbs and grasses for survival; insects serve as food sources for birds and their young and as forage for small and large herbivores.

Limy Residual ecological sites may be found in five plant community states (1.0 Reference State, 2.0 Native/Invaded State, 3.0 Invaded State, 4.0 Wooded Invaded, and 5.0 Go-Back) within a local landscape. Multiple plant community phases exist in states 1.0, 2.0 and 3.0. Today, these states occur primarily in response to grazing, drought, and non-use. Secondary influences include anthropogenic disturbances, black-tailed prairie dogs, and fire.

Because there is no known restoration pathway from State 2.0 to State 1.0, it is important to intensively manage using tools in State 1.0 and State 2.0 community phase pathways to prevent further plant community degradation along either the T2A transitional pathway to State 3.0. Native wildlife generally benefits from a heterogeneous composition and stature of grasslands found in States 1.0 and 2.0 which include diverse grass and forb species with varying stature and density. As plant communities degrade within State 2.0, warm-season grasses (particularly short-statured grasses) increase while native forbs are reduced. This transition results in reduced stature and increased plant community homogeneity. When adjacent/intermingled ecological sites undergo the same transition, the result can be an expansive, homogenous landscape.

Success along Restoration Pathway R3A from State 3.0 to State 2.0 is very difficult and is dependent upon presence of a remnant native grass population. This concept also applies to wildlife, as the target species must either be present on adjacent State 1.0 or State 2.0 plant communities or ecological sites within the species’ mobility limits. Species with limited mobility, such as Dakota skippers, must exist near the plant community to utilize restored sites. Mobile species (such as grassland-nesting birds) can more easily locate and utilize isolated, restored plant communities.

Plant community phases within the State 3.0 show dramatic increased homogeneity of exotic cool- season grasses, and further reduction in native forbs. Reduced forb diversity limits insect populations, negatively affecting grassland-nesting bird foraging opportunities. Increased exotic- grass litter can limit access to bare ground by nesting insects and can limit mobility by small chicks. A homogenous grassland landscape does not provide quality escape or winter cover. As a result, many species are not able to meet life requisites on communities in the 3.0 State.

Management along community phases, transition or restoration pathways should focus upon attainable changes. Short- and long-term monetary costs must be evaluated against short- and long- term ecological services in creating and maintaining habitat of sufficient quality to support a sustainable population.

1.0 Reference State

Community Phase 1.1 Western Wheatgrass-Needlegrasses-Blue Grama-Little Bluestem: This plant community offers quality vegetative cover for wildlife; every effort should be made to maintain this ecological site within this community phase. This phase retains high functionality through continued maintenance, including prescribed grazing with adequate recovery period as well as prescribed burning. Predominance of grass species in this community favors grazers and mixed-feeders (animals selecting grasses as well as forbs and shrubs). The structural diversity provides habitat for a wide array of migratory and resident birds. Calcium carbonate in the upper soil profile limits diversity of forbs.

Invertebrates: Insects play a role in maintaining the forb community and provide a forage base for grassland birds, reptiles, and rodents. Ecological services, historically provided by bison, are simulated by domestic livestock. These services include putting plant material and dung in contact with mineral soil to be used by low trophic-level consumers (such as invertebrate shredders, predators, herbivores, dung beetles, and fungal-feeders).

Dakota skippers may use this site due to presence of host plants, such as little bluestem and prairie dropseed. Regal fritillary habitat is limited due to the rarity of Nuttall's violet and prairie violets. Monarch butterfly may use flowering forbs on this site; however, few milkweed species are found on this site to support breeding. Bumblebees and other native bees utilize forbs as a nectar source and bare ground for nesting sites in bunchgrasses. Prescribed grazing with adequate recovery periods (as well as prescribed burning) to maintain the 1.1 phase has little effect on nests of ground-dwelling insects.

Birds: This plant community provides quality nesting, foraging, and escape habitats favored by short- to mid- grass-nesting birds. Plant stature may be too dense or tall for burrowing owl and McCown’s longspur; however, it may be used during periods of drought or management such as rotational grazing or fire, resulting in defoliation (along Community Phase Pathway 1.1A).

Several species of grassland birds that prefer mid-grass stature will use this site. In years with reduced precipitation or drought, use by mid-grass nesting species may be compromised. This plant community provides suitable areas for sharp-tailed grouse leks, nesting, and brood-rearing habitat. Limited stature and diverse prey populations provide good hunting opportunities for grassland raptors.

Mammals: The diversity of grasses and forbs provides high nutrition levels for small and large herbivores including voles, mice, rodents, jackrabbits, pronghorn, and deer (white-tailed and mule). Short to moderate stature provides suitable food and thermal, protective, and escape cover for small herbivores such as the hispid pocket mouse.

Amphibians/Reptiles: This ecological site and associated plant communities provide habitat for smooth green snakes. This ecological site can provide habitat for the northern leopard frog and Great Plains toad if freshwater habitats (such as stock water ponds) are located in or adjacent to the site. This ecological site provides limited habitat for the plains hog-nosed snake (prefer sandy soils) and plains spadefoot (prefer gravelly or sandy soils).

Fish and Mussels: This ecological site is not directly associated with streams, rivers, or water bodies. Associated ecological sites, such as Loamy Terrace, can receive run-on hydrology from Limy Residual sites. Management on these interconnected sites will have limited, secondary effects on aquatic species.

Community Phase 1.2 Blue Grama-Western Wheatgrass/Sedges/Forbs: Blue grama and western wheatgrass will dominate after multiyear drought with or without heavy long-term grazing; forb density, but not diversity, is slightly increased.

Invertebrates: Provides similar life requisites as Community Phase 1.1; however, long-term, heavy continuous grazing may negatively impact ground-nesting sites for bumble bees, other native bees, and other ground-nesting insects due to reduction of forbs, timing of forb flowering, or increased soil compaction.

Birds: This plant community provides quality nesting, foraging, and escape habitats favored by short grass-nesting birds. A shift to shorter plant stature along Community Phase Pathway 1.1A benefits McCown’s longspur, chestnut collared longspur, horned lark, and burrowing owl. Species preferring mid-grass stature will be generally successful with normal to above normal precipitation and a change in management along the 1.2A community phase pathway. In years with reduced precipitation or heavy grazing, nesting recruitment may be compromised for mid-grass-nesting species. This plant community provides areas suitable for sharp-tailed grouse leks. Limited cover and diverse prey populations provide good hunting opportunities for grassland raptors.

Mammals: Provides similar life requisites as Community Phase 1.1.

Amphibians/Reptiles: Provides similar life requisites as Community Phase 1.1.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

Community Phase 1.3 Little Bluestem-Western Wheatgrass/Sedges/Forbs: Physical soil surface disturbances, such as wind/water erosion and deposition in Community Phase Pathway 1.1B, cause increased calcium carbonates at the surface giving little bluestem a competitive advantage. This plant community occurs in a mosaic with Community Phase 1.1, creating a patchwork landscape. Little bluestem-dependent species, such as Dakota skipper, can utilize this habitat. Invertebrates: Provides similar life requisites as Community Phase 1.1.

Birds: Provides similar life requisites as Community Phase 1.1.

Mammals: Provides similar life requisites as Community Phase 1.1.

Amphibians/Reptiles: Provides similar life requisites as Community Phase 1.1.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

Community Phase 1.4 Annual Forbs/Bare Ground/Blue Grama/Fetid Marigold (Prairie Dog Town): This plant community phase is characterized by grazing-tolerant species and annual forbs (e.g., fetid marigold). Continued heavy grazing, repeated drought, prairie dog occupation, or a combination of these disturbances will shift to increased annual forbs with a reduction in perennial grasses. Perennial forbs stature and abundance are being replaced by short-statured annual forbs. Bare ground increases, litter amounts and infiltration rates decline while soil surface temperatures increase. Scruffy in appearance, this plant community is resilient and retains sufficient grazing sensitive native species to return to the 1.2 community phase (via Community Phase Pathway 1.4A).

Invertebrates: A switch to annual forbs from perennial forbs may have a significant impact to invertebrates, due to the loss of season-long nectar-producing plants for pollinators. Annual forbs’ bloom periods are dependent upon climatic conditions, especially precipitation events, and may not provide season-long nectaring opportunities. Season-long nectar sources may be found on adjacent plant communities or ecological sites for mobile species. Increased bare ground provides increased nesting sites for bumble bees and other ground-nesting insects.

Birds: This short-statured phase, driven by short-term prairie dog occupation, is favored by burrowing owls, chestnut-collared longspur, and McCown’s longspur. Prairie dog towns provide abundant prey populations for grassland raptors. The lack of grass and forb stature limits use by many bird species. Long-term prairie dog occupation following Transitional Phase Pathway T1B leads to the 2.0 State with no known return pathway due to the presence of exotic cool-season grasses. Managing this phase along Community Phase Pathway 1.4A can be an economical and successful method to restore habitat for many grassland-nesting birds.

Mammals: Suitable food, thermal, shelter, and escape cover (reduction in litter) for most mammals becomes limited. The loss of diversity of grasses and forbs reduces nutrition levels for small and large herbivores including rodents, white-tailed jackrabbits, and deer. Grazers, such as pronghorn, use prairie dog towns for foraging and loafing. Long-term prairie dog occupation following Community Phase Pathway T1B leads to the 2.0 State with no known return pathway due to the presence of exotic, cool-season grasses. Managing this phase along Community Phase Pathway 1.4A can be an economical and successful method to restore vegetative cover.

Amphibians/Reptiles: Prairie dog towns provide habitat for both amphibians and reptiles. Tiger salamanders, prairie rattlesnakes, and other snake species will use the burrow systems of prairie dogs for shelter and denning.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

2.0 Native/Invaded State

The main ecological driver for the Native/Invaded State is the invasion of exotic cool-season grasses. Limy Residual ecological sites have clumps or patches of naturally occurring buffaloberry and skunkbush sumac. Lack of fire can cause these small clumps to expand into larger clumps. Dominated by buffaloberry, these patches vary in size and are usually located on north and east facing slopes. Found on shoulder, back, or side slopes, a mosaic of grasses and shrubs may dominate the landscape on Limy Residual and associated Shallow Loamy ecological sites. These patches provide escape and thermal cover as well as food for many animals, including deer and sharp-tailed grouse. Loggerhead shrike are known to cache food by impaling prey animals on the long thorns. Bird species which are sensitive to woody species may avoid nesting in grassland areas with large amounts of woody vegetation.

Community Phase 2.1 Western Wheatgrass-Needlegrasses-Blue Grama-Little Bluestem /Shrubs: This plant community develops through Transition pathway T1A, due to changes in management and the presence of exotic, cool-season grasses. The threshold between states 1.0 and 2.0 is crossed when Kentucky bluegrass, crested wheatgrass, smooth brome, or other exotic species become established. This plant community phase has a very similar appearance and function to the Reference State of Community 1.1, except it has a minor amount of cool-season exotic grasses and forbs. This community phase is commonly found in mosaic in association with the 2.2 Community Phase. This phase functions at a high level for native wildlife; therefore, managers should consider the 2.0 Community Phase Pathways to avoid transitioning to State 3.0.

Invertebrates: Provides similar life requisites as Community Phase 1.1.

Birds: Provides similar life requisites as Community Phase 1.1.

Mammals: Provides similar life requisites as Community Phase 1.1.

Amphibians/Reptiles: Provides similar life requisites as Community Phase 1.1.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

Community Phase 2.2 Blue Grama-Western Wheatgrass-Little Bluestem/Sedges: Continuous, heavy season-long grazing or heavy seasonal grazing with or without drought (along Community Phase Pathway 2.1A) leads to shorter-statured species, such as blue grama and sedges. Dominated by shorter-stature grasses and a loss of nitrogen-fixing or leguminous native forbs, the diversity of this plant community is reduced. Both tap-rooted and fibrous-rooted perennial forbs increase in this phase but remain a minor component. Prescribed grazing with adequate recovery periods along Community Phase Pathway 2.2A is an efficient, effective method to regain the cool-season grass and forb diversity components in Community Phase 2.1.

Invertebrates: The loss of native forbs and increase in turf-forming grasses limit foraging and nesting sites for all pollinators. Continuous, heavy season-long grazing or heavy seasonal grazing may reduce ground-nesting site availability. Homogeneity of forb species may limit season long nectar availability.

Birds: Continuous, heavy season-long grazing or heavy seasonal grazing will reduce nesting sites, forage (invertebrates), and cover. A reduced forb component may limit foraging opportunities. Stature is generally short, serving both mid- and short-grass-nesting birds. Short-grass nesting birds favor this phase. Species preferring mid-grass stature will be generally successful with normal to above normal precipitation and a change in management along the 2.3A community phase pathway. In years with reduced precipitation or heavy grazing during the nesting season, nesting recruitment may be compromised for mid-grass-nesting species. This plant community provides areas suitable for sharp-tailed grouse lek site development. Limited stature and diverse prey populations provide good hunting opportunities for grassland raptors.

Mammals: Suitable food, thermal, protective, and escape cover (reduction in litter) for most mammals is somewhat reduced due to the increase in blue grama and sedges.

Amphibians/Reptiles: Provides similar life requisites as Community Phase 1.1.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

Community Phase 2.3 Blue grama/Sedges/Prairie Sagewort/Cactus: Continued heavy grazing with severe multiyear drought along Community Phase Pathway 2.2B leads to shorter-statured grasses (e.g., blue grama), sedges, shrubs and cacti. Plant community diversity is reduced with a loss of nitrogen-fixing or leguminous native forbs and minor grass components. Prescribed grazing with adequate recovery periods and return to normal precipitation along Community Phase Pathway 2.3A is the most effective method to regain diverse cool-season grass and forb components in Community Phase 2.2.
Insects: Provides similar life requisites as Community Phase 2.2.

Birds: Continued heavy grazing with severe multiyear drought, will reduce nesting sites, forage (insects and other invertebrates), and cover. Burrowing owl and McCown’s longspur rely on the stature and composition this plant community provides. This plant community provides areas suitable for sharp-tailed grouse leks. Diverse prey populations are available for grassland raptors. Management for bird species preferring mid-statured grasses should follow Community Phase Pathway 2.3A and 2.2A.

Mammals: Provides similar life requisites as Community Phase 2.2. The loss of diversity of grasses and forbs reduces nutrition levels for some small and large herbivores.

Amphibians/Reptiles: Provides similar life requisites as Community Phase 1.1.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

Community Phase 2.4 Annual Forbs/Bare Ground/Blue Grama/Fetid Marigold (Prairie Dog Town): This plant community is a result of ecological services provided by long-term, black-tailed prairie dog occupation (via Community Phases 2.1C, 2.2C, or 2.3B) coupled with the introduction of exotic cool- season grasses and annual forbs. Utilizing one or more tools in Community Phase Pathway 2.4A (e.g., removal of black-tailed prairie dogs, control of exotic perennial forbs, implementation of prescribed grazing) can move this community back to phase 2.1; but this may require significant management and economic inputs. Black-tailed prairie dogs provide primary ecological services to transition to and maintain Plant Community Phase 2.4.

Invertebrates: The loss of native forb diversity limits use by all pollinators. However, perennial invasive forbs will provide limited seasonal use dependent on bloom period. Bare ground, burrows, and short plant stature provide nest sites for bumblebees and other ground-nesting insects. Burrowing owls place dung around their burrow entrance, attracting dung beetles and other insects as a food source.

Birds: Burrowing owl and McCown’s longspur rely on the stature and composition that this plant community provides. Presence of black-tailed prairie dogs provided diverse prey populations for grassland raptors, including burrowing owls, prairie falcons, and ferruginous hawks. Burrowing owls nest in abandoned prairie dog burrows.

Mammals: Suitable food, thermal, protective, and escape cover (reduction in litter) for most mammals becomes limited. The loss of grass and forb diversity reduces nutrition levels for small and large herbivores including voles, mice, rodents, white-tailed jackrabbits, cottontail rabbits, and deer. Except for black-tailed prairie dog, this plant community provides little habitat for mid or small herbivores. Nonetheless, black-tailed prairie dog towns provide important habitat for many mammal species, including small rodents. Grazers, such as pronghorn, use prairie dog towns for foraging and loafing.

Amphibians/Reptiles: Prairie dog towns provide habitat for both amphibians and reptiles. Tiger salamanders, prairie rattlesnakes, and other snake species will use the burrow systems of prairie dogs for shelter and denning.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

Community Phase 2.5 Little Bluestem-Western Wheatgrass/Sedges/Forbs/Shrubs: This plant community develops through Community Phase Pathway 2.1b due to events (such as wind and/or water erosion, drought, pipeline and road installation and livestock trailing). These soil disturbances result in elevated calcium carbonates at the soil surface which favors an increase in little bluestem.

Invertebrates: Provides similar life requisites as Community Phase 1.1.

Birds: Provides similar life requisites as Community Phase 1.1.

Mammals: Provides similar life requisites as Community Phase 1.1.

Amphibians/Reptiles: Provides similar life requisites as Community Phase 1.1.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

3.0 Invaded State

Community Phase 3.1 Exotic Cool-Season Grasses/Forbs: Community Phase Pathway T2A is characterized by non-use or low intensity (<20% utilization) grazing and elimination of fire when exotic cool-season grasses are present as in Community Phase 2.0. This plant community phase is characterized by a dominance (>30%) of exotic cool-season grasses (such as Kentucky bluegrass, crested wheatgrass, and smooth brome). Restoration Pathway R3A requires remnant amounts of native warm-season grasses (i.e., blue grama, red threeawn, plains muhly), cool-season grasses (i.e., needlegrasses, western wheatgrass, prairie Junegrass), and forbs (i.e., white sagebrush, silverleaf Indian breadroot, blacksamson echinacea).

Frequent prescribed burns and high levels of grazing management along Restoration Pathway R3A are necessary to improve competitiveness and increase vigor and density of the remnant native plant community. The R3A pathway will have significant short-term negative impacts on wildlife cover. However, this is necessary to restore long-term native vegetation functions for various species’ habitat needs. Without intensive management, further degradation within State 3.0 will occur, requiring mechanical treatment with reseeding through Restoration Pathway R3B.

Invertebrates: Non-use or low intensity (<20% utilization) grazing, limits use by beneficial insects provided in States 1.0 and 2.0. Increased litter and lack of grazing leads to limited contact between plant material and mineral soil resulting in a cooler micro-climate, which is unfavorable to most insects. Lack of bare soil limits ground-nesting sites for native bees and other ground-nesting insects. The lack of nectar-producing plants limits forage opportunities for bumblebees, regal fritillary, monarch butterfly, and other pollinating species.

Birds: This homogeneous community phase, dominated by exotic plant species, provides limited vegetative cover and life requisites for most obligate grassland-nesting birds. Lack of stature and plant diversity, along with increased litter and the tendency of Kentucky bluegrass and smooth brome to lay down, limits use by many grassland-nesting birds. Litter accumulations reduce use by chestnut-collared and McCown’s longspurs. Burrowing owls may use the site if sufficient burrows of black-tailed prairie dogs or other burrowing mammals exist. Sharp-tailed grouse leks can be found on this plant community; however, winter cover must be provided by adjacent ecological sites or plant communities.

Mammals: Black-tailed prairie dog expansion is possible in this plant community phase. This community phase provides foraging habitat for pronghorn and deer. Litter accumulation favors thermal, protective, and escape cover for small rodents. However, reduced availability of native grass seed may reduce food availability for species such as the hispid pocket mouse.

Amphibians/Reptiles: Provides similar life requisites as Community Phase 1.1.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

4.0 Invaded Wooded State

Community Phase 4.1 Shrubs/Trees/Herbaceous: Resulting from pathways T2B and T3A, elimination of fire and grazing are the major contributors to this community phase crossing the threshold from an herbaceous plant community to a community dominated by shrubs and hardwoods. Limy Residual sites did not historically support a woody plant community. This community phase occupies north and east facing slopes.

Invertebrates: Dominated by early season flowering shrubs, pollinating insects may need adjacent herbaceous, and forb dominated ecological sites for mid- to late-season pollen sources. Lower trophic level consumers (such as invertebrate decomposers, scavengers, shredders, predators, herbivores, dung beetles, and fungal-feeders) will use woody plant material, leaves, and a limited number of grasses in contact with mineral soil. The woody component of this site is not conducive to use by the Dakota skipper, regal fritillary, or monarch butterfly. Woody plant material is available for wood nesting bees. These wind-protected, moist plant communities provide favorable habitat for flying insects (flies, mosquitoes, moths, etc.). Favorable climatic conditions can lead to large hatches of insects providing forage.

Birds: This site no longer provides habitat for grassland nesting bird species due to the woody vegetation dominance. Bird species that use and benefit from woodland edge (such as wild turkey, black-billed cuckoo, red-headed woodpecker, black-capped chickadee, gray catbird, and loggerhead shrike) can be found in this community phase. These sites provide sharp-tailed grouse brood cover and winter thermal cover. Wildlife use increases as the depth of snow increases during the winter, thereby becoming critical to the sustainment of winter resident bird populations. The presence of woody plant species may increase mammalian and avian predation and increase brood parasitism by brown-headed cowbirds on adjacent grassland ecological sites.

Mammals: Small herbivores that can use or tolerate woodland edge (such as least chipmunks, American porcupine, and cotton-tail rabbit) will benefit from this plant community phase. Shrubs and trees provide security and thermal cover used by elk and deer for foraging, loafing, and rearing young-of-the-year. Multi-layer shrub/tree communities provide concealment protection from predators during parturition. Plant species provide highly nutritious forage during peak lactation, one of the most energy demanding time periods of the year for female ungulates.

Deer utilize Community Phase 4.1 as primary foraging areas during all seasons of the year. Research at Theodore Roosevelt National Park found utilization of chokecherry and western snowberry by mule deer peaks during the fall accounting for 20 to 25% of their total dietary intake. In addition, mule deer also utilize rose, gooseberry, and serviceberry during the growing season. Winter white-tailed deer diets are dominated by chokecherry, western snowberry, Saskatoon serviceberry, rose, and various species of gooseberry. However, utilization of green ash as a forage by elk and deer was minimal accounting for <3% total dietary intake during all seasons.

Amphibians/Reptiles: Community Phase 4.1 will not support most of the amphibians and reptiles of conservation concern.

Fish and Mussels: Provides similar life requisites as Community Phase 1.1.

5.0 Go-Back State

Community Phase 5.1 Annual/Pioneer Perennial/Exotics: These plant communities are the result of severe soil disturbance (such as cropping, recreational activity, or concentrated livestock activity) for a prolonged period. Following cessation of disturbances, the resulting plant community is dominated by early pioneer annual and perennial plant species. Plant species composition and production are highly variable. Weedy plants can provide pollen sources along with spring and summer cover for many mammals and birds and their young. Dense weed cover can keep soils moist, increasing insect presence. Tall stature provided by some weeds, such as marsh elder and ragweed, offer thermal cover and seeds throughout winter.

Successful restoration of native species along Transition Pathway R5A results in a native grass and forb community in State 2.0. Failed restoration to native species through Restoration Pathway R5B results in Invaded State 3.0. Wildlife species response will be dependent upon plant community composition, vegetative stature, patch size, and management activities (such as prescribed grazing, burning, interseeding, haying, or noxious weed control).

Animal Community – Grazing Interpretations

This site is well adapted to managed grazing by domestic livestock. The predominance of herbaceous plants across all plant community phases best lends these sites to grazing by cattle, but other domestic grazers with differing diet preferences may also be a consideration depending upon management objectives. Often, the current plant community does not match any particular plant community (as described in the ecological site description). Because of this, a resource inventory is necessary to document plant composition and production. Proper interpretation of this inventory data will permit the establishment of a safe, initial stocking rate for the type and class of animals and level of grazing management. More accurate stocking rate estimates should eventually be calculated using actual stocking rate information and monitoring data.

NRCS defines prescribed grazing as “managing the harvest of vegetation with grazing and/or browsing animals with the intent to achieve specific ecological, economic, and management objectives”. As used in this site description, the term ‘prescribed grazing’ is intended to include multiple grazing management systems (e.g., rotational grazing, twice-over grazing, conservation grazing, targeted grazing, etc.) provided that, whatever management system is implemented, it meets the intent of prescribed grazing definition.

The basic grazing prescription addresses balancing forage demand (quality and quantity) with available forage, varying grazing and deferment periods from year-to-year, matching recovery/deferment periods to growing conditions when pastures are grazed more than once in a growing season, implementation of a contingency (e.g., drought) plan, and a monitoring plan. When the management goal is to facilitate change from one plant community phase or state to another, then the prescription needs to be designed to shift the competitive advantage to favor the native grass and forb species.

Grazing levels are noted within the plant community narratives and pathways in reference to grazing management. “Degree of utilization” is defined as the proportion of the current years forage production that is consumed and/or destroyed by grazing animals (may refer to a single plant species or a portion or all the vegetation). “Grazing utilization” is classified as slight, moderate, full, close, and severe (see the following table for description of each grazing use category). The following utilization levels are also described in the Ranchers Guide to Grassland Management IV. Utilization levels are determined by using the landscape appearance method as outlined in the Interagency Technical Reference “Utilization Studies and Residual Measurements” 1734-3.

Utilization Level % Use Description
Slight (Light) 0-20 Appears practically undisturbed when viewed obliquely. Only choice areas and forage utilized.

Moderate 20-40 Almost all of accessible range shows grazing. Little or no use of poor forage. Little evidence of trailing to grazing.

Full 40-60 All fully accessible areas are grazed. The major sites have key forage species properly utilized (about half taken, half left). Points of concentration with overuse limited to 5 to 10 percent of accessible area.

Close (Heavy) 60-80 All accessible range plainly shows use and major sections closely cropped. Livestock forced to use less desirable forage, considering seasonal preference.

Severe > 80 Key forage species completely used. Low-value forages are dominant.

Hydrological functions

Water is the principal factor limiting herbage production on this site. The site is dominated by soils in hydrologic groups B and C. Infiltration rate is moderate and runoff potential for this site varies from low to high depending upon hydrologic group, slope, and ground cover. In many cases, areas with greater than 75% ground cover have the greatest potential for high infiltration and lower runoff. An exception would be where short grasses form a dense sod and dominate the site. Areas where ground cover is less than 50% have the greatest potential to have reduced infiltration and higher runoff (refer to Section 4, NRCS National Engineering Handbook for runoff quantities and hydrologic curves).

Recreational uses

The largest acreage of public land available for recreation in the MLRA is owned and managed by the United States Forest Service (USFS) within the Little Missouri, Grand River, and Cedar River National Grasslands in South Dakota and the Little Missouri National Grasslands in North Dakota (687,398 acres). These areas are available for hunting, fishing, hiking, camping, horse and bike riding, nature viewing, etc. In addition, the Bureau of Land Management (BLM) manages (40,264 acres) in North and South Dakota with the same recreational opportunities as the USFS lands.

The United States Army Corps of Engineers (USAE) owns 496,162 acres of land and water located on and adjacent to Lake Sakakawea and Lake Oahe. The North Dakota and South Dakota Game and Fish Departments manage the fisheries resources. These two Missouri River reservoirs provide excellent fishing and water recreation opportunities. In addition, the United States Fish and Wildlife Service (USFWS) manages a national fish hatchery below Garrison Dam.

The USFWS manages 36,858 acres in the National Wildlife Refuge system while the North Dakota and South Dakota wildlife management agencies manage 72,218 acres as wildlife or game management areas. The North Dakota, South Dakota, and Montana Department of Trust Lands manage 486,482 acres. These areas provide hunting, bird watching, hiking, and other outdoor recreation opportunities. North Dakota Wildlife Management Areas along the shoreline of Lake Sakakawea and the Missouri River account for 60,000 acres of the approximately 72,218 acres of land managed by the states for wildlife habitat in MLRA 54. Located in the northern portion of the MLRA, the Killdeer Mountain WMA is the largest tract of state-owned land managed for wildlife habitat at approximately 7,000 acres.

The largest refuge managed by the United States Fish and Wildlife service is Lake Ilo National Wildlife Refuge totaling approximately 4,000 acres. United States Bureau of Reclamation manages approximately 11,000 acres at Lake Tschida and 8,460 acres at Bowman-Haley Lake for fish and wildlife habitat. The National Park Service manages the Knife River Indian Village National Historic Site; the North Dakota Historical Society manages the Double Ditch Indian Village site.

Bird watching: Public and private grasslands within MLRA 54 provide essential habitat for prairie-dependent bird species such as Sprague's pipits, western meadowlark, and Baird's sparrow along with some of the larger, showy members of the upland prairie include marbled godwits, upland sandpipers, willets, and sharp-tailed grouse. Publicly owned lands provide excellent birding opportunities. MLRA 54 is in the Central Flyway.

Hunting/Fishing: MLRA 54 is a fall destination for thousands of pheasant and upland game bird hunters. This MLRA also provides excellent deer (white-tailed and mule), pronghorn, and coyote hunting opportunities. Lake Sakakawea, Lake Oahe, Lake Tschida, and the Missouri River provide excellent year-round fishing opportunities. The North Dakota Game and Fish Department and South Dakota Game, Fish and Parks manage approximately 40 fishing lakes within the MLRA. Available species include yellow perch, walleye, northern pike, muskellunge, crappie, bluegill, rainbow trout, and smallmouth bass. Chinook salmon are stocked in Lake Sakakawea.

Camping: Numerous state operated campgrounds are located along the shores of Lake Sakakawea, Lake Oahe, Missouri River, and Shadehill Reservoir. Primitive camping is allowed on Grand River and Cedar River National Grasslands in South Dakota and the Little Missouri National Grasslands in North Dakota. Other numerous camping (primitive and improved) sites are available in numerous city and county parks.

Hiking/Biking/Horseback Riding: Hiking is permitted on most state and federally owned lands. Developed hiking and biking trails can be found on Harmon Lake (13.1 miles), Roughrider Trail (Morton County, 16.5 miles), Missouri River State Natural Area (5 miles), Ft. Abraham Lincoln State Park (8 miles), Cross Ranch State Park (14 miles), Grand River National Grasslands (7 miles), Lake Sakakawea State Park (5 miles), and Lewis & Clark State Park (5 miles). In addition, extensive biking and walking trails are found in local county and city parks. Ft. Abraham Lincoln State Park has 6 miles of horseback trails.

Wood products

No appreciable wood products are present on the site.

Other products

Seed harvest of native plant species can provide additional income on this site.

Other information

Site Development and Testing Plan
• NASIS revisions needed:
o Consider relinking Lantry components from Thin Loamy to Limy Residual.
This ESD is the best available current knowledge. The site concept and species composition table have been used in the field and tested for more than five years. It is expected that as additional information becomes available revisions may be required.

Inventory data references

Information presented here has been derived from NRCS and other federal/state agency clipping and inventory data. Also, field knowledge of range-trained personnel was used. All descriptions were peer reviewed and/or field-tested by various private, state, and federal agency specialists.

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Bluemle. J.P. 2016. North Dakota’s geologic legacy. North Dakota State University Press. 382 pages.

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Contributors

ND NRCS: David Dewald, Jonathan Fettig, Jody Forman, Mike Gerbig, Alan Gulsvig, Mark Hayek, Jeanne Heilig, John Kempenich, Chuck Lura, Jeff Printz, Steve Sieler, and Hal Weiser.

Approval

Suzanne Mayne-Kinney, 3/31/2025

Acknowledgments

NRCS would like to acknowledge the United State Forest Service (USFS) and National Park Service (NPS) for access to USFS properties and technical assistance in ESD development. USFS: Jack Dahl, Nickole Dahl, and Chad Prosser.